About the Authors

Ilan Ben-Zvi

Affiliation Division of Autoimmune and Musculoskeletal Disease, Feinstein Institute for Medical Research, Manhasset, New York, United States of America

Cynthia Aranow

Affiliation Division of Autoimmune and Musculoskeletal Disease, Feinstein Institute for Medical Research, Manhasset, New York, United States of America

Meggan Mackay

Affiliation Division of Autoimmune and Musculoskeletal Disease, Feinstein Institute for Medical Research, Manhasset, New York, United States of America

Anfisa Stanevsky

Affiliation Division of Autoimmune and Musculoskeletal Disease, Feinstein Institute for Medical Research, Manhasset, New York, United States of America

Diane L. Kamen

Affiliation Division of Rheumatology, Medical University of South Carolina, Charleston, South Carolina, United States of America

L. Manuela Marinescu

Affiliation Department of Medicine, Albert Einstein College of Medicine, The Bronx, New York, United States of America

Christopher E. Collins

Affiliation Department of Medicine, Ralph H. Johnson Veterans Administration Medical Center, Charleston, South Carolina, United States of America

Gary S. Gilkeson

Affiliations Division of Rheumatology, Medical University of South Carolina, Charleston, South Carolina, United States of America, Department of Medicine, Ralph H. Johnson Veterans Administration Medical Center, Charleston, South Carolina, United States of America

Betty Diamond

Affiliation Division of Autoimmune and Musculoskeletal Disease, Feinstein Institute for Medical Research, Manhasset, New York, United States of America

John A. Hardin

johnhardin@arthritis.org

Affiliations Department of Medicine, Albert Einstein College of Medicine, The Bronx, New York, United States of America, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Washington, D.C., United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CA BD JH. Performed the experiments: IBZ AS JH. Analyzed the data: IBZ CA MM AS LMM GSG BD JH. Contributed reagents/materials/analysis tools: IBZ CA MM DLK LMM CEC GSG. Wrote the paper: IBZ CA MM BD JH.