About the Authors

Pawandeep Dhami

Current address: University College London Cancer Institute, University College London, London, United Kingdom

Affiliation The Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Alexander W. Bruce

Current address: Wellcome Trust/Cancer Research UK Gurdon Institute, University of Cambridge, Cambridge, United Kingdom

Affiliation The Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Johanna H. Jim

Current address: Section of Haemato-Oncology, Institute of Cancer Research, Sutton, United Kingdom

Affiliation The Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Shane C. Dillon

Current address: School of Genetics and Microbiology, Moyne Institute of Preventative Medicine, Trinity College, Dublin, Ireland

Affiliation The Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Amanda Hall

Affiliation The Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Jonathan L. Cooper

Affiliation The Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Nicolas Bonhoure

Affiliation Section of Pathology and Gene Regulation, Division of Cancer Sciences and Molecular Pathology, University of Glasgow, Glasgow, United Kingdom

Kelly Chiang

Affiliation Section of Pathology and Gene Regulation, Division of Cancer Sciences and Molecular Pathology, University of Glasgow, Glasgow, United Kingdom

Peter D. Ellis

Affiliation The Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Cordelia Langford

Affiliation The Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Robert M. Andrews

Affiliation The Wellcome Trust Sanger Institute, Hinxton, United Kingdom

David Vetrie

dlv1q@clinmed.gla.ac.uk

Affiliations The Wellcome Trust Sanger Institute, Hinxton, United Kingdom, Section of Pathology and Gene Regulation, Division of Cancer Sciences and Molecular Pathology, University of Glasgow, Glasgow, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PD AWB JJ SCD DV. Performed the experiments: PD AWB JJ SCD AH JLC NB KC. Analyzed the data: PD AWB JJ SCD AH JLC NB KC RMA DV. Contributed reagents/materials/analysis tools: PE CL RMA. Wrote the paper: DV.