About the Authors

Barend Mark de Clare Bronsvoort

Mark.Bronsvoort@ed.ac.uk

Affiliation The Roslin Institute and The Royal (Dick) School of Veterinary Studies, University of Edinburgh, Roslin, Midlothian, United Kingdom

Beatrix von Wissmann

Affiliation Centre for Infectious Diseases, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

Eric Maurice Fèvre

Affiliation Centre for Infectious Diseases and Centre for Infection, Immunity and Evolution, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Ian Graham Handel

Affiliation The Roslin Institute and The Royal (Dick) School of Veterinary Studies, University of Edinburgh, Roslin, Midlothian, United Kingdom

Kim Picozzi

Affiliation Centre for Infectious Diseases, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

Sue Christina Welburn

Affiliation Centre for Infectious Diseases, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BMdCB BvW EMF KP SCW. Performed the experiments: BvW KP. Analyzed the data: BMdCB IGH. Wrote the paper: BMdCB BvW EMF IGH SCW.