About the Authors

Nobuo Imai

inobuo@kais.kyoto-u.ac.jp

Affiliation Graduates School of Agriculture, Kyoto University, Sakyo-ku, Kyoto, Japan

Hiromitsu Samejima

Affiliation Graduates School of Agriculture, Kyoto University, Sakyo-ku, Kyoto, Japan

Andreas Langner

Affiliation Graduates School of Agriculture, Kyoto University, Sakyo-ku, Kyoto, Japan

Robert C. Ong

Affiliation Forest Research Centre, Sabah Forestry Department, Sandakan, Sabah, Malaysia

Satoshi Kita

Affiliation Graduates School of Agriculture, Kyoto University, Sakyo-ku, Kyoto, Japan

Jupiri Titin

Affiliation Forest Research Centre, Sabah Forestry Department, Sandakan, Sabah, Malaysia

Arthur Y. C. Chung

Affiliation Forest Research Centre, Sabah Forestry Department, Sandakan, Sabah, Malaysia

Peter Lagan

Affiliation Deramakot Forest Office, Sabah Forestry Department, Sandakan, Sabah, Malaysia

Ying Fah Lee

Affiliation Forest Research Centre, Sabah Forestry Department, Sandakan, Sabah, Malaysia

Kanehiro Kitayama

Affiliation Graduates School of Agriculture, Kyoto University, Sakyo-ku, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NI HS AL RCO SK. Performed the experiments: NI HS AL RCO SK JT. Analyzed the data: NI HS AL RCO SK JT. Contributed reagents/materials/analysis tools: AYCC PL YFL KK. Wrote the paper: NI HS AL RCO SK JT AYCC PL YFL KK.