About the Authors

Melissa A. Edeling

Contributed equally to this work with: Melissa A. Edeling, Subramaniam Sanker

Affiliation Cambridge Institute for Medical Research, University of Cambridge, Cambridge, United Kingdom

Subramaniam Sanker

Contributed equally to this work with: Melissa A. Edeling, Subramaniam Sanker

Affiliation Department of Cell Biology and Physiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Takaki Shima

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

P. K. Umasankar

Affiliation Department of Cell Biology and Physiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Stefan Höning

Affiliation Institute of Biochemistry I and Center for Molecular Medicine, University of Cologne, Cologne, Germany

Hye Y. Kim

Affiliation Department of Bioengineering, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Lance A. Davidson

Affiliation Department of Bioengineering, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Simon C. Watkins

Affiliation Department of Cell Biology and Physiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Michael Tsang

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

David J. Owen

Affiliation Cambridge Institute for Medical Research, University of Cambridge, Cambridge, United Kingdom

Linton M. Traub

traub@pitt.edu

Affiliation Department of Cell Biology and Physiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MAE SS TS PKU SH HYK LAD MT DJO LMT. Performed the experiments: MAE SS TS PKU SH HYK SCW MT DJO LMT. Analyzed the data: MAE SS TS PKU SH HYK LAD SCW MT DJO LMT. Contributed reagents/materials/analysis tools: LAD SCW MT DJO LMT. Wrote the paper: MAE DJO LMT. Contributed to preparation of the manuscript: SS PKU LAD MT.