About the Authors

Daniel N. Frank

Affiliations Department of Molecular, Cellular, and Developmental Biology, University of Colorado, Boulder, Colorado, United States of America, Mucosal and Vaccine Research Program Colorado, University of Colorado, Denver, Colorado, United States of America

Shandra S. Wilson

Affiliation Department of Urologic Oncology, University of Colorado Denver and Health Sciences Center, Aurora, Colorado, United States of America

Allison L. St. Amand

Affiliation Department of Molecular, Cellular, and Developmental Biology, University of Colorado, Boulder, Colorado, United States of America

Norman R. Pace

norman.pace@colorado.edu

Affiliation Department of Molecular, Cellular, and Developmental Biology, University of Colorado, Boulder, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DF SSW NP. Performed the experiments: DF ALSA. Analyzed the data: DF ALSA NP. Contributed reagents/materials/analysis tools: DF SSW. Wrote the paper: DF NP.