About the Authors

Renata I. Mazzucchelli

Current address: Laboratory of Gene Therapy and Primary Immunodeficiency, San Raffaele Telethon Institute for Gene Therapy, HSR-TIGET, Milano, Italy,

Affiliation Laboratory of Molecular Immunoregulation, Cancer and Inflammation Program, Center for Cancer Research, National Cancer Institute, National Institute of Health, Frederick, Maryland, United States of America

Søren Warming

Current address: Genentech, Inc., Department of Molecular Biology, South San Francisco, California, United States of America

Affiliation Mouse Cancer Genetics Program, National Cancer Institute, Frederick, Maryland, United States of America

Scott M. Lawrence

Affiliation Laboratory Animal Science Program (LASP), Science Applications International Corporation (SAIC), Cancer Research Center, National Cancer Institute, Frederick, Maryland, United States of America

Masaru Ishii

Current address: Laboratory of Biological Imaging, Immunology Frontier Research Center, Osaka University, Osaka, Japan

Affiliation Lymphocyte Biology Section, Laboratory of Immunology, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Mehrnoosh Abshari

Current address: Twinbrook Flow Cytometry Facility, Lockheed Martin Contractor, RTB/NIAID/NIH, Rockville, Maryland, United States of America

Affiliation Laboratory Animal Science Program (LASP), Science Applications International Corporation (SAIC), Cancer Research Center, National Cancer Institute, Frederick, Maryland, United States of America

A. Valance Washington

Current address: Department of Biology, University of Puerto Rico Recinto Mayaguez, Mayaguez, Puerto Rico

Affiliation Laboratory of Experimental Immunology, Cancer and Inflammation Program, Center for Cancer Research, National Cancer Institute, National Institute of Health, Frederick, Maryland, United States of America

Lionel Feigenbaum

Affiliation Laboratory Animal Science Program (LASP), Science Applications International Corporation (SAIC), Cancer Research Center, National Cancer Institute, Frederick, Maryland, United States of America

Andrew C. Warner

Affiliation Laboratory Animal Science Program (LASP), Science Applications International Corporation (SAIC), Cancer Research Center, National Cancer Institute, Frederick, Maryland, United States of America

Davis J. Sims

Affiliation Laboratory Animal Science Program (LASP), Science Applications International Corporation (SAIC), Cancer Research Center, National Cancer Institute, Frederick, Maryland, United States of America

Wen Qing Li

Affiliation Laboratory of Molecular Immunoregulation, Cancer and Inflammation Program, Center for Cancer Research, National Cancer Institute, National Institute of Health, Frederick, Maryland, United States of America

Julie A. Hixon

Affiliation Laboratory of Molecular Immunoregulation, Cancer and Inflammation Program, Center for Cancer Research, National Cancer Institute, National Institute of Health, Frederick, Maryland, United States of America

Daniel H. D. Gray

Affiliation The Walter and Eliza Hall Institute of Medical Research, Parkville, Victoria, Australia

Benjamin E. Rich

Affiliation Department of Dermatology, Brigham and Women's Hospital, Boston, Massachusetts, United States of America

Matthew Morrow

Affiliation Human Retrovirus Section, Vaccine Branch, Center for Cancer Research NCI, Frederick, Maryland, United States of America

Miriam R. Anver

Affiliation Laboratory Animal Science Program (LASP), Science Applications International Corporation (SAIC), Cancer Research Center, National Cancer Institute, Frederick, Maryland, United States of America

James Cherry

Affiliation Gene Expression Laboratory, SAIC-Frederick, Inc., NCI-Frederick, Frederick, Maryland, United States of America

Dieter Naf

Affiliation Laboratory Animal Science Program (LASP), Science Applications International Corporation (SAIC), Cancer Research Center, National Cancer Institute, Frederick, Maryland, United States of America

Lawrence R. Sternberg

Affiliation Laboratory Animal Science Program (LASP), Science Applications International Corporation (SAIC), Cancer Research Center, National Cancer Institute, Frederick, Maryland, United States of America

Daniel W. McVicar

Affiliation Laboratory of Experimental Immunology, Cancer and Inflammation Program, Center for Cancer Research, National Cancer Institute, National Institute of Health, Frederick, Maryland, United States of America

Andrew G. Farr

Affiliation Department of Immunology, University of Washington, Seattle, Washington, United States of America

Ronald N. Germain

Affiliation Lymphocyte Biology Section, Laboratory of Immunology, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Keith Rogers

Affiliation Laboratory Animal Science Program (LASP), Science Applications International Corporation (SAIC), Cancer Research Center, National Cancer Institute, Frederick, Maryland, United States of America

Nancy A. Jenkins

Current address: Institute of Molecular and Cell Biology, Proteos, Singapore

Affiliation Mouse Cancer Genetics Program, National Cancer Institute, Frederick, Maryland, United States of America

Neal G. Copeland

Current address: Institute of Molecular and Cell Biology, Proteos, Singapore

Affiliation Mouse Cancer Genetics Program, National Cancer Institute, Frederick, Maryland, United States of America

Scott K. Durum

durums@ncifcrf.gov

Affiliation Laboratory of Molecular Immunoregulation, Cancer and Inflammation Program, Center for Cancer Research, National Cancer Institute, National Institute of Health, Frederick, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RIM AVW DWM RNG NAJ NGC SKD. Performed the experiments: RIM SW SML MI MA LF ACW DJS JAH MM JC DN LRS AGF KR. Analyzed the data: RIM WQL MRA AGF SKD. Contributed reagents/materials/analysis tools: DHDG BER. Wrote the paper: RIM SKD.