About the Authors

Sandra R. Schulze

Affiliation Department of Biology, Western Washington University, Bellingham, Washington, United States of America

Beatrice Curio-Penny

Affiliation Department of Biochemistry, University of Iowa, Iowa City, Iowa, United States of America

Sean Speese

Affiliation Department of Neurobiology, University of Massachusetts, Wochester, Massachusetts, United States of America

George Dialynas

Affiliation Department of Biochemistry, University of Iowa, Iowa City, Iowa, United States of America

Diane E. Cryderman

Affiliation Department of Biochemistry, University of Iowa, Iowa City, Iowa, United States of America

Caitrin W. McDonough

Affiliation Department of Biochemistry, University of Iowa, Iowa City, Iowa, United States of America

Demet Nalbant

Affiliation Department of Biochemistry, University of Iowa, Iowa City, Iowa, United States of America

Melissa Petersen

Affiliation Department of Biology, Western Washington University, Bellingham, Washington, United States of America

Vivian Budnik

Affiliation Department of Neurobiology, University of Massachusetts, Wochester, Massachusetts, United States of America

Pamela K. Geyer

Affiliation Department of Biochemistry, University of Iowa, Iowa City, Iowa, United States of America

Lori L. Wallrath

lori-wallrath@uiowa.edu

Affiliation Department of Biochemistry, University of Iowa, Iowa City, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SRS SS GD LLW. Performed the experiments: SRS BCP SS GD DEC CWM DN MP. Analyzed the data: SRS BCP SS GD DEC CWM DN MP VB PKG LLW. Wrote the paper: SRS LLW.