About the Authors

Holly E. Copeland

hcopeland@tnc.org

Affiliation The Nature Conservancy, Lander, Wyoming, United States of America

Kevin E. Doherty

Affiliation National Audubon Society, Laramie, Wyoming, United States of America

David E. Naugle

Affiliation Wildlife Biology Program, University of Montana, Missoula, Montana, United States of America

Amy Pocewicz

Affiliation The Nature Conservancy, Lander, Wyoming, United States of America

Joseph M. Kiesecker

Affiliation The Nature Conservancy, Lander, Wyoming, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HEC KED DEN AP JMK. Analyzed the data: HEC KED. Contributed reagents/materials/analysis tools: HEC. Wrote the paper: HEC DEN AP JMK.