About the Authors

Lora L. Shadwick

lora.lindley@gmail.com

Affiliation Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas, United States of America

Frederick W. Spiegel

Affiliation Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas, United States of America

John D. L. Shadwick

Affiliation Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas, United States of America

Matthew W. Brown

Affiliation Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas, United States of America

Jeffrey D. Silberman

Affiliation Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LLS FWS JDS. Performed the experiments: LLS JDLS MWB. Analyzed the data: LLS JDLS JDS. Contributed reagents/materials/analysis tools: FWS JDS. Wrote the paper: LLS FWS JDLS JDS.