About the Authors

Robert K. Bradley

Affiliation Biophysics Graduate Group, University of California, Berkeley, California, United States of America

Andrew V. Uzilov

Affiliation Department of Bioengineering, University of California, Berkeley, California, United States of America

Mitchell E. Skinner

Affiliation Department of Bioengineering, University of California, Berkeley, California, United States of America

Yuri R. Bendaña

Affiliation Department of Bioengineering, University of California, Berkeley, California, United States of America

Lars Barquist

Affiliation Department of Bioengineering, University of California, Berkeley, California, United States of America

Ian Holmes

ihh@berkeley.edu

Affiliations Biophysics Graduate Group, University of California, Berkeley, California, United States of America, Department of Bioengineering, University of California, Berkeley, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RKB IH. Performed the experiments: RKB AVU MS YB LB IH. Analyzed the data: RKB AVU MS YB LB IH. Contributed reagents/materials/analysis tools: RKB AVU MS IH. Wrote the paper: RKB IH.