About the Authors

Yuya Makiguchi

Affiliation Division of Biosphere Science, Graduate School of Environmental Science, Hokkaido University, Sapporo, Hokkaido, Japan

Shinya Nagata

Affiliation Dainippon Sumitomo Pharma Co. Ltd., Suita, Osaka, Japan

Takahito Kojima

Affiliation College of Bioresource Sciences, Nihon University, Fujisawa, Kanagawa, Japan

Masaki Ichimura

Affiliation Shibetsu Salmon Museum, Shibetsu, Hokkaido, Japan

Yoshifumi Konno

Affiliation Division of Biosphere Science, Graduate School of Environmental Science, Hokkaido University, Sapporo, Hokkaido, Japan

Hideki Murata

Affiliation Dainippon Sumitomo Pharma Co. Ltd., Suita, Osaka, Japan

Hiroshi Ueda

hueda@fsc.hokudai.ac.jp

Affiliations Division of Biosphere Science, Graduate School of Environmental Science, Hokkaido University, Sapporo, Hokkaido, Japan, Field Science Center for Northern Biosphere, Hokkaido University, Sapporo, Hokkaido, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YM SN MI. Performed the experiments: YM MI YK. Analyzed the data: YM SN TK HU. Contributed reagents/materials/analysis tools: SN TK HM. Wrote the paper: YM HU.