About the Authors

Ayumi Kishioka

Affiliation Department of Molecular Neurobiology and Pharmacology, Graduate School of Medicine, University of Tokyo, Tokyo, Japan

Fumiaki Fukushima

Affiliation Department of Molecular Neurobiology and Pharmacology, Graduate School of Medicine, University of Tokyo, Tokyo, Japan

Tamae Ito

Affiliation Department of Molecular Neurobiology and Pharmacology, Graduate School of Medicine, University of Tokyo, Tokyo, Japan

Hirotaka Kataoka

Affiliation Department of Molecular Neurobiology and Pharmacology, Graduate School of Medicine, University of Tokyo, Tokyo, Japan

Hisashi Mori

Current address: Department of Molecular Neuroscience, Graduate School of Medicine and Pharmaceutical Sciences, University of Toyama, Toyama, Japan

Affiliation Department of Molecular Neurobiology and Pharmacology, Graduate School of Medicine, University of Tokyo, Tokyo, Japan

Toshio Ikeda

Current address: Laboratory of Experimental Animal Model Research, National Center for Geriatrics and Gerontology, Obu, Japan

Affiliation Laboratory of Behavioral Genetics, Brain Science Institute, RIKEN, Saitama, Japan

Shigeyoshi Itohara

Affiliation Laboratory of Behavioral Genetics, Brain Science Institute, RIKEN, Saitama, Japan

Kenji Sakimura

Affiliation Department of Cellular Neurobiology, Brain Research Institute, Niigata University, Niigata, Japan

Masayoshi Mishina

mishina@m.u-tokyo.ac.jp

Affiliation Department of Molecular Neurobiology and Pharmacology, Graduate School of Medicine, University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MM. Performed the experiments: AK FF TI HK. Analyzed the data: AK FF TI. Contributed reagents/materials/analysis tools: HK HM TI SI KS. Wrote the paper: AK FF MM.