About the Authors

Shin-ichi Hashimoto

Contributed equally to this work with: Shin-ichi Hashimoto, Wei Qu

Affiliation Department of Molecular Preventive Medicine, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Wei Qu

Contributed equally to this work with: Shin-ichi Hashimoto, Wei Qu

Affiliation Department of Computational Biology, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwa, Japan

Budrul Ahsan

Affiliation Department of Computational Biology, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwa, Japan

Katsumi Ogoshi

Affiliation Department of Molecular Preventive Medicine, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Atsushi Sasaki

Affiliation Department of Computational Biology, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwa, Japan

Yoichiro Nakatani

Affiliation Department of Computational Biology, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwa, Japan

Yongjun Lee

Affiliation Department of Molecular Preventive Medicine, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Masako Ogawa

Affiliation Department of Molecular Preventive Medicine, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Akio Ametani

Affiliation Department of Molecular Preventive Medicine, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Yutaka Suzuki

Affiliation Department of Medical Genome Science, Graduate School of Frontier Science, The University of Tokyo, Kashiwa, Japan

Sumio Sugano

Affiliation Department of Medical Genome Science, Graduate School of Frontier Science, The University of Tokyo, Kashiwa, Japan

Clarence C. Lee

Affiliation Applied Biosystems, Foster City, California, United States of America

Robert C. Nutter

Affiliation Applied Biosystems, Foster City, California, United States of America

Shinichi Morishita

Affiliation Department of Computational Biology, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwa, Japan

Kouji Matsushima

koujim@m.u-tokyo.ac.jp

Affiliation Department of Molecular Preventive Medicine, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SiH WQ. Performed the experiments: SiH KO YL MO YS CCL RCN. Analyzed the data: SiH WQ BA AS YN SM. Contributed reagents/materials/analysis tools: SiH WQ YS SS. Wrote the paper: SiH WQ AA SS SM KM.