About the Authors

Saori Morino

Contributed equally to this work with: Saori Morino, Tatsuya Kondo

Affiliation Department of Molecular Medicine, Faculty of Medical and Pharmaceutical Sciences, Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Tatsuya Kondo

Contributed equally to this work with: Saori Morino, Tatsuya Kondo

Affiliation Department of Metabolic Medicine, Faculty of Medical and Pharmaceutical Sciences, Kumamoto University, Kumamoto, Japan

Kazunari Sasaki

Affiliation Department of Metabolic Medicine, Faculty of Medical and Pharmaceutical Sciences, Kumamoto University, Kumamoto, Japan

Hironori Adachi

Affiliation Department of Metabolic Medicine, Faculty of Medical and Pharmaceutical Sciences, Kumamoto University, Kumamoto, Japan

Mary Ann Suico

Affiliation Department of Molecular Medicine, Faculty of Medical and Pharmaceutical Sciences, Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Erika Sekimoto

Affiliation Department of Molecular Medicine, Faculty of Medical and Pharmaceutical Sciences, Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Tomoko Matsuda

Affiliation Department of Molecular Medicine, Faculty of Medical and Pharmaceutical Sciences, Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Tsuyoshi Shuto

Affiliation Department of Molecular Medicine, Faculty of Medical and Pharmaceutical Sciences, Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Eiichi Araki

earaki@gpo.kumamoto-u.ac.jp (EA); hirokai@gpo.kumamoto-u.ac.jp (HK)

Affiliation Department of Metabolic Medicine, Faculty of Medical and Pharmaceutical Sciences, Kumamoto University, Kumamoto, Japan

Hirofumi Kai

earaki@gpo.kumamoto-u.ac.jp (EA); hirokai@gpo.kumamoto-u.ac.jp (HK)

Affiliation Department of Molecular Medicine, Faculty of Medical and Pharmaceutical Sciences, Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SM TK EA HK. Performed the experiments: SM TK KS HA ES TM. Analyzed the data: SM TK EA HK. Contributed reagents/materials/analysis tools: TS EA HK. Wrote the paper: SM TK MAS.