About the Authors

Hiroaki Kimura

Affiliation Department of Orthopaedics, Kyoto University, Kyoto, Japan

Kin Ming Kwan

Affiliations Department of Molecular Genetics, The University of Texas M. D. Anderson Cancer Center, Houston, Texas, United States of America, Department of Biology, The Chinese University of Hong Kong, Shatin, Hong Kong SAR, China

Zhaoping Zhang

Affiliation Department of Molecular Genetics, The University of Texas M. D. Anderson Cancer Center, Houston, Texas, United States of America

Jian Min Deng

Affiliation Department of Molecular Genetics, The University of Texas M. D. Anderson Cancer Center, Houston, Texas, United States of America

Bryant G. Darnay

Affiliation Department of Experimental Therapeutics, The University of Texas M. D. Anderson Cancer Center, Houston, Texas, United States of America

Richard R. Behringer

Affiliation Department of Molecular Genetics, The University of Texas M. D. Anderson Cancer Center, Houston, Texas, United States of America

Takashi Nakamura

Affiliation Department of Orthopaedics, Kyoto University, Kyoto, Japan

Benoit de Crombrugghe

Affiliation Department of Molecular Genetics, The University of Texas M. D. Anderson Cancer Center, Houston, Texas, United States of America

Haruhiko Akiyama

hakiyama@kuhp.kyoto-u.ac.jp

Affiliation Department of Orthopaedics, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BdC HA. Performed the experiments: HK KMK BGD HA. Analyzed the data: HK BGD RRB TN HA. Contributed reagents/materials/analysis tools: ZZ JMD RRB BdC. Wrote the paper: HK BdC HA.