About the Authors

Linda Munson

Affiliation Department of Pathology, Microbiology and Immunology, University of California Davis, Davis, California, United States of America

Karen A. Terio

Affiliation University of Illinois Zoological Pathology Program, Maywood, Illinois, United States of America

Richard Kock

Affiliation African Union International Bureau for Animal Resources, Nairobi, Kenya

Titus Mlengeya

Affiliation Tanzania National Parks, Arusha, Tanzania

Melody E. Roelke

Affiliations Tanzania National Parks, Arusha, Tanzania, Laboratory of Genomic Diversity, SAIC-Frederick, Inc., NCI-Frederick, Frederick, Maryland, United States of America

Edward Dubovi

Affiliation Department of Population Medicine and Diagnostic Sciences, Cornell University, Ithaca, New York, United States of America

Brian Summers

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Anthony R. E. Sinclair

Affiliation Department of Zoology, University of British Columbia, Vancouver, Canada

Craig Packer


Affiliation Department of Ecology, Evolution, and Behavior, University of Minnesota, St. Paul, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CP LM KT. Performed the experiments: LM KT ED BS. Analyzed the data: TS CP RK LM KT ED BS. Contributed reagents/materials/analysis tools: TS CP TM RK MR. Wrote the paper: TS CP RK LM KT BS.