About the Authors

Elizabeth A. Reynolds Losin

Affiliations Division of Psychobiology, Yerkes National Primate Research Center, Atlanta, Georgia, United States of America, Interdepartmental Program in Neuroscience, University of California Los Angeles, Los Angeles, California, United States of America

Jamie L. Russell

Affiliation Division of Psychobiology, Yerkes National Primate Research Center, Atlanta, Georgia, United States of America

Hani Freeman

Affiliation Division of Psychobiology, Yerkes National Primate Research Center, Atlanta, Georgia, United States of America

Adrien Meguerditchian

Affiliation Division of Psychobiology, Yerkes National Primate Research Center, Atlanta, Georgia, United States of America

William D. Hopkins

whopkin@emory.edu

Affiliations Division of Psychobiology, Yerkes National Primate Research Center, Atlanta, Georgia, United States of America, Department of Psychology, Agnes Scott College, Decatur, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WH ER. Performed the experiments: WH JR HF ER. Analyzed the data: WH AM ER. Wrote the paper: WH ER.