About the Authors

Jung Yoon Park

Affiliations Department of Medicine, Albert Einstein College of Medicine, Bronx, New York, United States of America, Department of Molecular Genetics, Albert Einstein College of Medicine, Bronx, New York, United States of America

Mi-Ook Cho

Affiliations Department of Medicine, Albert Einstein College of Medicine, Bronx, New York, United States of America, Department of Molecular Genetics, Albert Einstein College of Medicine, Bronx, New York, United States of America

Shanique Leonard

Affiliation Department of Physiology, Barshop Institute for Longevity and Aging Studies, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America

Brent Calder

Affiliation Buck Institute for Age Research, Novato, California, United States of America

I. Saira Mian

Affiliation Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Woo Ho Kim

Affiliation Department of Pathology, Seoul National University College of Medicine, Seoul, Korea

Susan Wijnhoven

Affiliation National Institute of Public Health and the Environment, Laboratory of Toxicology, Pathology and Genetics, Bilthoven, the Netherlands

Harry van Steeg

Affiliation National Institute of Public Health and the Environment, Laboratory of Toxicology, Pathology and Genetics, Bilthoven, the Netherlands

James Mitchell

Affiliation MGC-Department of Cell Biology and Genetics, Erasmus University Medical Center, Rotterdam, the Netherlands

Gijsbertus T. J. van der Horst

Affiliation MGC-Department of Cell Biology and Genetics, Erasmus University Medical Center, Rotterdam, the Netherlands

Jan Hoeijmakers

Affiliation MGC-Department of Cell Biology and Genetics, Erasmus University Medical Center, Rotterdam, the Netherlands

Pinchas Cohen

Affiliation Pediatric Endocrinology, University of California Los Angeles, Los Angeles, California, United States of America

Jan Vijg

Affiliation Buck Institute for Age Research, Novato, California, United States of America

Yousin Suh

ysuh@aecom.yu.edu

Affiliations Department of Medicine, Albert Einstein College of Medicine, Bronx, New York, United States of America, Department of Molecular Genetics, Albert Einstein College of Medicine, Bronx, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JV YS. Performed the experiments: PC JP MC SL WK. Analyzed the data: IM RC. Contributed reagents/materials/analysis tools: JH Gv JM Hv SW. Wrote the paper: JV YS.