About the Authors

Chee Gee Liew

Current address: UCI Stem Cell Research Center, University of California Irvine, Irvine, California, United States of America

Affiliations Centre for Stem Cell Biology, Department of Biomedical Science, The University of Sheffield, Sheffield, United Kingdom, Centre for Stem Cell Biology, Department of Molecular Biology and Biotechnology, The University of Sheffield, Sheffield, United Kingdom

Nadia N. Shah

Contributed equally to this work with: Nadia N. Shah, Sarah J. Briston

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Sarah J. Briston

Contributed equally to this work with: Nadia N. Shah, Sarah J. Briston

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Ruth M. Shepherd

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Cheen Peen Khoo

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Mark J. Dunne

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Harry D. Moore

Affiliations Centre for Stem Cell Biology, Department of Biomedical Science, The University of Sheffield, Sheffield, United Kingdom, Centre for Stem Cell Biology, Department of Molecular Biology and Biotechnology, The University of Sheffield, Sheffield, United Kingdom

Karen E. Cosgrove

karen.e.cosgrove@manchester.ac.uk (KC); p.w.andrews@sheffield.ac.uk (PA)

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Peter W. Andrews

karen.e.cosgrove@manchester.ac.uk (KC); p.w.andrews@sheffield.ac.uk (PA)

Affiliations Centre for Stem Cell Biology, Department of Biomedical Science, The University of Sheffield, Sheffield, United Kingdom, Centre for Stem Cell Biology, Department of Molecular Biology and Biotechnology, The University of Sheffield, Sheffield, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PA CL KC. Performed the experiments: CL NS SB RS CK. Analyzed the data: PA CL NS SB RS CK MD KC. Contributed reagents/materials/analysis tools: PA HM RS MD KC. Wrote the paper: PA HM CL MD KC. Other: Designed the study: PA.