About the Authors

Ashok Hegde

Affiliation Laboratory of Developmental Signalling and Patterning, Institute of Molecular and Cell Biology, A*STAR (Agency for Science, Technology and Research), Singapore, Singapore

Nick Chuanxin Qiu

Contributed equally to this work with: Nick Chuanxin Qiu, Xuehui Qiu, Steven Hao-Kee Ho, Kenny Qi-Ye Tay

Affiliation Laboratory of Developmental Signalling and Patterning, Institute of Molecular and Cell Biology, A*STAR (Agency for Science, Technology and Research), Singapore, Singapore

Xuehui Qiu

Contributed equally to this work with: Nick Chuanxin Qiu, Xuehui Qiu, Steven Hao-Kee Ho, Kenny Qi-Ye Tay

Affiliation Laboratory of Developmental Signalling and Patterning, Institute of Molecular and Cell Biology, A*STAR (Agency for Science, Technology and Research), Singapore, Singapore

Steven Hao-Kee Ho

Contributed equally to this work with: Nick Chuanxin Qiu, Xuehui Qiu, Steven Hao-Kee Ho, Kenny Qi-Ye Tay

Affiliation Laboratory of Developmental Signalling and Patterning, Institute of Molecular and Cell Biology, A*STAR (Agency for Science, Technology and Research), Singapore, Singapore

Kenny Qi-Ye Tay

Contributed equally to this work with: Nick Chuanxin Qiu, Xuehui Qiu, Steven Hao-Kee Ho, Kenny Qi-Ye Tay

Affiliation Laboratory of Developmental Signalling and Patterning, Institute of Molecular and Cell Biology, A*STAR (Agency for Science, Technology and Research), Singapore, Singapore

Joshy George

Affiliation Genome Institute of Singapore, A*STAR (Agency for Science, Technology and Research), Singapore, Singapore

Felicia Soo Lee Ng

Affiliation Bioinformatics Institute, A*STAR (Agency for Science, Technology and Research), Singapore, Singapore

Kunde Ramamoorthy Govindarajan

Affiliation Genome Institute of Singapore, A*STAR (Agency for Science, Technology and Research), Singapore, Singapore

Zhiyuan Gong

Affiliation Department of Biological Sciences, National University of Singapore, Singapore, Singapore

Sinnakaruppan Mathavan

Affiliation Genome Institute of Singapore, A*STAR (Agency for Science, Technology and Research), Singapore, Singapore

Yun-Jin Jiang

To whom correspondence should be addressed. E-mail: yjjiang@imcb.a-star.edu.sg

Affiliations Laboratory of Developmental Signalling and Patterning, Institute of Molecular and Cell Biology, A*STAR (Agency for Science, Technology and Research), Singapore, Singapore, Department of Biochemistry, National University of Singapore, Singapore, Singapore, School of Biological Sciences, Nanyang Technological University, Singapore, Singapore

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YJ AH. Performed the experiments: AH KT NQ SH XQ. Analyzed the data: KG YJ JG AH FN SM. Contributed reagents/materials/analysis tools: ZG YJ SM. Wrote the paper: YJ AH SM.