About the Authors

Saori C. Tanaka

To whom correspondence should be addressed. E-mail: xsaori@atr.jp (ST); doya@oist.jp (KD)

Affiliations Department of Computational Neurobiology, ATR Computational Neuroscience Laboratories, Seika, Souraku, Kyoto, Japan, Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Seika, Souraku, Kyoto, Japan

Nicolas Schweighofer

Affiliations Department of Computational Neurobiology, ATR Computational Neuroscience Laboratories, Seika, Souraku, Kyoto, Japan, Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Seika, Souraku, Kyoto, Japan, Department of Biokinesiology and Physical Therapy, University of Southern California, Los Angeles, California, United States of America

Shuji Asahi

Affiliations Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Seika, Souraku, Kyoto, Japan, Department of Psychiatry and Neurosciences, Hiroshima University, Minamiku, Hiroshima, Japan

Kazuhiro Shishida

Affiliations Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Seika, Souraku, Kyoto, Japan, Department of Psychiatry and Neurosciences, Hiroshima University, Minamiku, Hiroshima, Japan

Yasumasa Okamoto

Affiliations Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Seika, Souraku, Kyoto, Japan, Department of Psychiatry and Neurosciences, Hiroshima University, Minamiku, Hiroshima, Japan

Shigeto Yamawaki

Affiliations Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Seika, Souraku, Kyoto, Japan, Department of Psychiatry and Neurosciences, Hiroshima University, Minamiku, Hiroshima, Japan

Kenji Doya

To whom correspondence should be addressed. E-mail: xsaori@atr.jp (ST); doya@oist.jp (KD)

Affiliations Department of Computational Neurobiology, ATR Computational Neuroscience Laboratories, Seika, Souraku, Kyoto, Japan, Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Seika, Souraku, Kyoto, Japan, Neural Computational Unit, Okinawa Institute of Science and Technology, Suzaki, Uruma, Okinawa, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NS YO ST SA SY KD. Performed the experiments: NS KS YO ST SA. Analyzed the data: NS ST. Wrote the paper: ST KD.