About the Authors

Kenji Matsuura

To whom correspondence should be addressed. E-mail: kenjijpn@cc.okayama-u.ac.jp

Affiliations Graduate School of Environmental Science, Okayama University, Okayama, Japan, Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America

Takashi Tamura

Affiliation Graduate School of Natural Science and Technology, Okayama University, Okayama, Japan

Norimasa Kobayashi

Affiliation Graduate School of Environmental Science, Okayama University, Okayama, Japan

Toshihisa Yashiro

Affiliation Graduate School of Environmental Science, Okayama University, Okayama, Japan

Shingo Tatsumi

Affiliation Graduate School of Environmental Science, Okayama University, Okayama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KM TT. Performed the experiments: KM TY NK ST. Analyzed the data: KM NK. Contributed reagents/materials/analysis tools: TT. Wrote the paper: KM.