About the Authors

Takumi Yamagishi

Affiliation Kondo Research Unit, Brain Development Research Group, Brain Science Institute, Institute of Physical and Chemical Research (RIKEN), Wako, Japan

Michiru Ozawa

Affiliation Kondo Research Unit, Brain Development Research Group, Brain Science Institute, Institute of Physical and Chemical Research (RIKEN), Wako, Japan

Chiho Ohtsuka

Affiliation Kondo Research Unit, Brain Development Research Group, Brain Science Institute, Institute of Physical and Chemical Research (RIKEN), Wako, Japan

Ritsuko Ohyama-Goto

Affiliation Kondo Research Unit, Brain Development Research Group, Brain Science Institute, Institute of Physical and Chemical Research (RIKEN), Wako, Japan

Takashi Kondo

To whom correspondence should be addressed. E-mail: TKondo@brain.riken.jp

Affiliation Kondo Research Unit, Brain Development Research Group, Brain Science Institute, Institute of Physical and Chemical Research (RIKEN), Wako, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK. Performed the experiments: TK MO TY CO RO. Analyzed the data: TK. Contributed reagents/materials/analysis tools: TK MO TY CO RO. Wrote the paper: TK.