About the Authors

Sarah L. Clarke

Contributed equally to this work with: Sarah L. Clarke, Gareth J. Betts, Awen M. Gallimore, Andrew J. Godkin

To whom correspondence should be addressed. E-mail: godkinaj@cf.ac.uk

Affiliation Department of Medical Biochemistry and Immunology, Cardiff University, Cardiff, United Kingdom

Gareth J. Betts

Contributed equally to this work with: Sarah L. Clarke, Gareth J. Betts, Awen M. Gallimore, Andrew J. Godkin

Affiliation Department of Medical Biochemistry and Immunology, Cardiff University, Cardiff, United Kingdom

Andrea Plant

Affiliation Department of Medical Biochemistry and Immunology, Cardiff University, Cardiff, United Kingdom

Kate L. Wright

Affiliation Department of Medical Biochemistry and Immunology, Cardiff University, Cardiff, United Kingdom

Tariq M. El-Shanawany

Affiliation Department of Medical Biochemistry and Immunology, Cardiff University, Cardiff, United Kingdom

Richard Harrop

Affiliation Oxford Biomedica, Oxford, United Kingdom

Jared Torkington

Affiliation Department of Surgery, University Hospital of Wales, Cardiff, United Kingdom

Brian I. Rees

Affiliation Department of Surgery, University Hospital of Wales, Cardiff, United Kingdom

Geraint T. Williams

Affiliation Department of Pathology, Cardiff University, Cardiff, United Kingdom

Awen M. Gallimore

Contributed equally to this work with: Sarah L. Clarke, Gareth J. Betts, Awen M. Gallimore, Andrew J. Godkin

Affiliation Department of Medical Biochemistry and Immunology, Cardiff University, Cardiff, United Kingdom

Andrew J. Godkin

Contributed equally to this work with: Sarah L. Clarke, Gareth J. Betts, Awen M. Gallimore, Andrew J. Godkin

Affiliation Department of Medical Biochemistry and Immunology, Cardiff University, Cardiff, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AG AG SC AP. Performed the experiments: SC GB AP KW TE. Analyzed the data: AG SC. Contributed reagents/materials/analysis tools: RH JT BR GW. Wrote the paper: AG AG SC. Other: Collected the data: SC GB AP KW TE. Collected human samples: JT BR GW.