About the Authors

Guillaume Fournié

Affiliations Galápagos Genetics Epidemiology and Pathology Laboratory, Galápagos National Park, Puerto Ayora, Santa Cruz, Galápagos Islands, Ecuador, Ecole Nationale Vétérinaire d’Alfort, Maisons-Alfort, France, Department of Production and Population Health, Royal Veterinary College, Hatfield, United Kingdom

Simon J. Goodman

Affiliations Galápagos Genetics Epidemiology and Pathology Laboratory, Galápagos National Park, Puerto Ayora, Santa Cruz, Galápagos Islands, Ecuador, School of Biology, Faculty of Biological Sciences, University of Leeds, Leeds, United Kingdom

Marilyn Cruz

Affiliation Galápagos Genetics Epidemiology and Pathology Laboratory, Galápagos National Park, Puerto Ayora, Santa Cruz, Galápagos Islands, Ecuador

Virna Cedeño

Affiliations Galápagos Genetics Epidemiology and Pathology Laboratory, Galápagos National Park, Puerto Ayora, Santa Cruz, Galápagos Islands, Ecuador, Concepto Azul, Guayaquil, Ecuador, Biotechnology Program, University of Guayaquil, Guayaquil, Ecuador

Alberto Vélez

Affiliation Galápagos Genetics Epidemiology and Pathology Laboratory, Galápagos National Park, Puerto Ayora, Santa Cruz, Galápagos Islands, Ecuador

Leandro Patiño

Affiliations Galápagos Genetics Epidemiology and Pathology Laboratory, Galápagos National Park, Puerto Ayora, Santa Cruz, Galápagos Islands, Ecuador, School of Biology, Faculty of Biological Sciences, University of Leeds, Leeds, United Kingdom, Institute of Zoology, Zoological Society of London, London, United Kingdom

Caroline Millins

Current address: Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Affiliations Galápagos Genetics Epidemiology and Pathology Laboratory, Galápagos National Park, Puerto Ayora, Santa Cruz, Galápagos Islands, Ecuador, Institute of Zoology, Zoological Society of London, London, United Kingdom

Lynda M. Gibbons

Affiliation Department of Pathology and Infectious Disease, Royal Veterinary College, London, United Kingdom

Mark T. Fox

Affiliation Department of Pathology and Infectious Disease, Royal Veterinary College, London, United Kingdom

Andrew A. Cunningham

a.cunningham@ioz.ac.uk

Affiliations Galápagos Genetics Epidemiology and Pathology Laboratory, Galápagos National Park, Puerto Ayora, Santa Cruz, Galápagos Islands, Ecuador, Institute of Zoology, Zoological Society of London, London, United Kingdom

Competing Interests

One of the authors, Virna Cedeno, is employed by a commercial company. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AAC SJG VC. Performed the experiments: GF MC AV LP CM LMG MTF. Analyzed the data: GF SJG. Contributed reagents/materials/analysis tools: AAC SJG VC. Wrote the paper: GF SJG AAC.