About the Authors

Georgios Banos

Affiliations SRUC, Roslin Institute Building, Edinburgh, United Kingdom, Faculty of Veterinary Medicine, Aristotle University of Thessaloniki, Thessaloniki, Greece

Eileen Wall

Affiliation SRUC, Roslin Institute Building, Edinburgh, United Kingdom

Michael P. Coffey

Affiliation SRUC, Roslin Institute Building, Edinburgh, United Kingdom

Ainsley Bagnall

Affiliation SRUC, Roslin Institute Building, Edinburgh, United Kingdom

Sandra Gillespie

Affiliation Moredun Research Institute, Edinburgh, United Kingdom

George C. Russell

Affiliation Moredun Research Institute, Edinburgh, United Kingdom

Tom N. McNeilly

Tom.McNeilly@moredun.ac.uk

Affiliation Moredun Research Institute, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GB EW MPC GR TNM. Performed the experiments: AB SG GR TNM. Analyzed the data: GB EW MPC. Contributed reagents/materials/analysis tools: AB. Wrote the paper: GB EW MPC AB SG GR TNM.