About the Authors

Ryutaro Goto

gotoryutaro@gmail.com

Affiliations Department of Marine Ecosystem Dynamics, Atmosphere and Ocean Research Institute, The University of Tokyo, Kashiwa, Chiba, Japan, Graduate School of Human and Environmental Studies, Kyoto University, Kyoto, Japan

Tomoko Okamoto

Affiliation Graduate School of Human and Environmental Studies, Kyoto University, Kyoto, Japan

Hiroshi Ishikawa

Affiliation Uwajima, Ehime, Japan

Yoichi Hamamura

Affiliation Kure, Hiroshima, Japan

Makoto Kato

Affiliation Graduate School of Human and Environmental Studies, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RG MK. Performed the experiments: RG. Analyzed the data: RG. Contributed reagents/materials/analysis tools: RG TO HI YH MK. Wrote the paper: RG TO MK.