About the Authors

Yoshinori Yamakawa

Affiliations Graduate School of Informatics, Kyoto University, Sakyo-ku, Kyoto, Japan, The Japan Society for the Promotion of Science, Tokyo, Japan, ATR Computational Neuroscience Laboratories, Kyoto, Japan, NEC Corporation Corporate Planning Division, Tokyo, Japan

Ryota Kanai

Affiliation Institute of Cognitive Neuroscience, University College London, London, United Kingdom

Michikazu Matsumura

Affiliation Graduate School of Human and Environmental Studies, Kyoto University, Sakyo-ku, Kyoto, Japan

Eiichi Naito

enaito@atr.jp

Affiliations ATR Computational Neuroscience Laboratories, Kyoto, Japan, National Institute of Information and Communication Technology, Research Department 1, Kobe Advanced ICT Research Center, Biophysical ICT Group, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YY EN. Performed the experiments: YY EN. Analyzed the data: YY RK EN. Contributed reagents/materials/analysis tools: YY EN. Wrote the paper: YY RK MM EN.