About the Authors

Ye Liu

Contributed equally to this work with: Ye Liu, He-Ping Zhang, Shou-Feng Zhang, Jin-Xiang Wang

Affiliation Laboratory of Epidemiology and Key Laboratory of Jilin Provincial Zoonosis Control and Prevention, Military Veterinary Research Institute, Academy of Military Medical Sciences, Changchun, China

He-Ping Zhang

Contributed equally to this work with: Ye Liu, He-Ping Zhang, Shou-Feng Zhang, Jin-Xiang Wang

Affiliation Ningxia Hui Autonomous Region Center for Animal Disease Control and Prevention, Yinchuan, China

Shou-Feng Zhang

Contributed equally to this work with: Ye Liu, He-Ping Zhang, Shou-Feng Zhang, Jin-Xiang Wang

Affiliation Laboratory of Epidemiology and Key Laboratory of Jilin Provincial Zoonosis Control and Prevention, Military Veterinary Research Institute, Academy of Military Medical Sciences, Changchun, China

Jin-Xiang Wang

Contributed equally to this work with: Ye Liu, He-Ping Zhang, Shou-Feng Zhang, Jin-Xiang Wang

Affiliation Ningxia Hui Autonomous Region Center for Animal Disease Control and Prevention, Yinchuan, China

Hai-Ning Zhou

Affiliation Ningxia Hui Autonomous Region Center for Animal Disease Control and Prevention, Yinchuan, China

Fei Zhang

Affiliation Laboratory of Epidemiology and Key Laboratory of Jilin Provincial Zoonosis Control and Prevention, Military Veterinary Research Institute, Academy of Military Medical Sciences, Changchun, China

Yu-Mei Wang

Affiliation Ningxia Hui Autonomous Region Center for Animal Disease Control and Prevention, Yinchuan, China

Long Ma

Affiliation Ningxia Hui Autonomous Region Center for Animal Disease Control and Prevention, Yinchuan, China

Nan Li

Affiliation Laboratory of Epidemiology and Key Laboratory of Jilin Provincial Zoonosis Control and Prevention, Military Veterinary Research Institute, Academy of Military Medical Sciences, Changchun, China

Rong-Liang Hu

ronglianghu@hotmail.com

Affiliation Laboratory of Epidemiology and Key Laboratory of Jilin Provincial Zoonosis Control and Prevention, Military Veterinary Research Institute, Academy of Military Medical Sciences, Changchun, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YL HPZ RLH. Performed the experiments: YL SFZ JXW HNZ YMW LM FZ NL. Analyzed the data: YL RLH. Contributed reagents/materials/analysis tools: HPZ JXW. Wrote the paper: YL RLH.