About the Authors

Ram K. Raghavan

rkraghavan@vet.k-state.edu

Affiliation Kansas State Veterinary Diagnostic Laboratory and Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Cathleen A. Hanlon

‡ CAH, DGG, and GAA also contributed equally to this work.

Affiliation Centers for Disease Control and Prevention, Atlanta, Georgia, United States of America

Douglas G. Goodin

‡ CAH, DGG, and GAA also contributed equally to this work.

Affiliation Department of Geography, College of Arts and Sciences, Kansas State University, Manhattan, Kansas, United States of America

Rolan Davis

Contributed equally to this work with: Rolan Davis, Michael Moore, Susan Moore

Affiliation Kansas State Veterinary Diagnostic Laboratory and Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Michael Moore

Contributed equally to this work with: Rolan Davis, Michael Moore, Susan Moore

Affiliation Kansas State Veterinary Diagnostic Laboratory and Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Susan Moore

Contributed equally to this work with: Rolan Davis, Michael Moore, Susan Moore

Affiliation Kansas State Veterinary Diagnostic Laboratory and Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Gary A. Anderson

‡ CAH, DGG, and GAA also contributed equally to this work.

Affiliation Kansas State Veterinary Diagnostic Laboratory and Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Competing Interests

The authors declare that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RKR CAH GAA. Performed the experiments: RKR DGG CAH. Analyzed the data: RKR DGG. Contributed reagents/materials/analysis tools: RKR MM RD SM. Wrote the paper: RKR CAH DGG RD SM GAA.