About the Authors

Catherine Grant


Affiliation Institute of Development Studies, Falmer, Brighton, United Kingdom

Neil Anderson

Affiliation The Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Roslin, United Kingdom

Noreen Machila

Affiliation School of Veterinary Medicine, University of Zambia, Lusaka, Zambia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CG NA NM. Performed the experiments: CG NM. Analyzed the data: CG. Contributed reagents/materials/analysis tools: CG NA NM. Wrote the paper: CG NA NM.