About the Authors

Kate M. Mitchell

Kate.Mitchell@lshtm.ac.uk

Current address: Department of Global Health and Development, London School of Hygiene and Tropical Medicine, London, United Kingdom

Affiliation Centre for Immunity, Infection and Evolution, Institute of Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Francisca Mutapi

Affiliation Centre for Immunity, Infection and Evolution, Institute of Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Takafira Mduluza

Affiliations Department of Biochemistry, University of Zimbabwe, Harare, Zimbabwe, College of Health Sciences, University of KwaZulu Natal, Durban, South Africa

Nicholas Midzi

Current address: Department of Medical Microbiology, University of Zimbabwe, Harare, Zimbabwe

Affiliation National Institute of Health Research, Harare, Zimbabwe

Nicholas J. Savill

Affiliation Centre for Immunity, Infection and Evolution, Institute of Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Mark E. J. Woolhouse

Affiliation Centre for Immunity, Infection and Evolution, Institute of Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KMM FM MEJW. Performed the experiments: KMM. Analyzed the data: KMM. Wrote the paper: KMM FM MEJW. Helped to conceive the research project: TM NM. Collected field data informing the modelling process: TM NM. Developed computer code: NJS. Reviewed the draft manuscript: NJS TM.