About the Authors

Hideo Hasegawa

Affiliation Department of Biology, Oita University, Yufu, Oita, Japan

David Modrý

Affiliations Department of Pathology and Parasitology, Faculty of Veterinary Medicine, University of Veterinary and Pharmaceutical Sciences, Brno, Czech Republic, Central European Institute of Technology, University of Veterinary and Pharmaceutical Sciences, Brno, Czech Republic, Biology Centre, Institute of Parasitology, Academy of Sciences of the Czech Republic, Ceske Budejovice, Czech Republic

Masahiro Kitagawa

Affiliation Department of Biology, Oita University, Yufu, Oita, Japan

Kathryn A. Shutt

Affiliation Department of Anthropology, Durham University, Durham, United Kingdom

Angelique Todd

Affiliation World Wildlife Foundation (WWF), Dzanga Sangha Protected Areas, Bangui, Central African Republic

Barbora Kalousová

Affiliation Department of Pathology and Parasitology, Faculty of Veterinary Medicine, University of Veterinary and Pharmaceutical Sciences, Brno, Czech Republic

Ilona Profousová

Affiliation Department of Pathology and Parasitology, Faculty of Veterinary Medicine, University of Veterinary and Pharmaceutical Sciences, Brno, Czech Republic

Klára J. Petrželková

petrzelkova@ivb.cz

Affiliations Department of Pathology and Parasitology, Faculty of Veterinary Medicine, University of Veterinary and Pharmaceutical Sciences, Brno, Czech Republic, Biology Centre, Institute of Parasitology, Academy of Sciences of the Czech Republic, Ceske Budejovice, Czech Republic, Liberec Zoo, Liberec, Czech Republic, Institute of Vertebrate Biology, Academy of Sciences of the Czech Republic, Brno, Czech Republic

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HH DM KJP. Performed the experiments: BK KAS IP DM AT KJP HH MK. Analyzed the data: HH DM MK KJP. Contributed reagents/materials/analysis tools: HH DM KAS KJP. Wrote the paper: HH BK KAS IP DM AT KJP.