About the Authors

Joe Kgaladi

Affiliation Department of Microbiology and Plant Pathology, Faculty of Natural and Agricultural Sciences, University of Pretoria, Pretoria, South Africa

Nicolette Wright

Affiliation Department of Microbiology and Plant Pathology, Faculty of Natural and Agricultural Sciences, University of Pretoria, Pretoria, South Africa

Jessica Coertse

Affiliation Department of Microbiology and Plant Pathology, Faculty of Natural and Agricultural Sciences, University of Pretoria, Pretoria, South Africa

Wanda Markotter

Affiliation Department of Microbiology and Plant Pathology, Faculty of Natural and Agricultural Sciences, University of Pretoria, Pretoria, South Africa

Denise Marston

Affiliation Wildlife Zoonoses and Vector-borne Diseases Research Group, OIE Rabies Reference Laboratory/WHO Collaborating Centre for the Characterization of Rabies and Rabies-related Viruses), Department of Virology, Animal Health Veterinary Laboratories Agency (Weybridge), Addlestone, Surrey, United Kingdom

Anthony R. Fooks

Affiliations Wildlife Zoonoses and Vector-borne Diseases Research Group, OIE Rabies Reference Laboratory/WHO Collaborating Centre for the Characterization of Rabies and Rabies-related Viruses), Department of Virology, Animal Health Veterinary Laboratories Agency (Weybridge), Addlestone, Surrey, United Kingdom, University of Liverpool, Department of Clinical Infection, Microbiology and Immunology, Liverpool, United Kingdom

Conrad M. Freuling

Affiliation OIE Rabies Reference Laboratory/WHO Collaborating Centre for Rabies Surveillance and Research, Friedrich-Loeffler-Institut, Federal Research Institute for Animal Health, Institute of Molecular Biology, Greifswald-Insel Riems, Germany

Thomas F. Müller

Thomas.Mueller@fli.bund.de

Affiliation OIE Rabies Reference Laboratory/WHO Collaborating Centre for Rabies Surveillance and Research, Friedrich-Loeffler-Institut, Federal Research Institute for Animal Health, Institute of Molecular Biology, Greifswald-Insel Riems, Germany

Claude T. Sabeta

Affiliation OIE Rabies Reference Laboratory, Agricultural Research Council, Onderstepoort Veterinary Institute, Pretoria, South Africa

Louis H. Nel

Affiliation Department of Microbiology and Plant Pathology, Faculty of Natural and Agricultural Sciences, University of Pretoria, Pretoria, South Africa

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LHN TFM ARF CTS. Performed the experiments: JK NW JC CMF DM WM CTS. Analyzed the data: JK NW JC CMF DM WM CTS. Contributed reagents/materials/analysis tools: JK NW JC WM DM ARF CFM TFM CTS LHN. Wrote the paper: JK CMF DM LHN TFM ARF CTS WM.