About the Authors

Beatrix von Wissmann

Affiliation Centre for Infectious Diseases, School of Biomedical Sciences, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom

Noreen Machila

Affiliation Centre for Infectious Diseases, School of Biomedical Sciences, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom

Kim Picozzi

Affiliation Centre for Infectious Diseases, School of Biomedical Sciences, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom

Eric M. Fèvre

Affiliation School of Biological Sciences, College of Science and Engineering, The University of Edinburgh, Edinburgh, United Kingdom

Barend M. deC. Bronsvoort

Affiliation Epidemiology, Economics and Risk Assessment (EERA), The Roslin Institute and Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Roslin, United Kingdom

Ian G. Handel

Affiliation Epidemiology, Economics and Risk Assessment (EERA), The Roslin Institute and Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Roslin, United Kingdom

Susan C. Welburn

sue.welburn@ed.ac.uk

Affiliation Centre for Infectious Diseases, School of Biomedical Sciences, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BvW NM KP EMF SCW. Performed the experiments: BvW KP SCW. Analyzed the data: BvW NM KP EMF BMdB IGH SCW. Contributed reagents/materials/analysis tools: SCW. Wrote the paper: BvW NM KP EMF BMdB SCW. Field work: SCW BvW NM KP EMF.