About the Authors

Nicholas A. Ettinger

Affiliations Interdisciplinary Graduate Program in Molecular and Cellular Biology, University of Iowa, Iowa City, Iowa, United States of America, Medical Scientist Training Program, University of Iowa Carver College of Medicine, Iowa City, Iowa, United States of America

Mary E. Wilson

mary-wilson@uiowa.edu

Affiliations Interdisciplinary Graduate Program in Molecular and Cellular Biology, University of Iowa, Iowa City, Iowa, United States of America, Medical Scientist Training Program, University of Iowa Carver College of Medicine, Iowa City, Iowa, United States of America, Departments of Internal Medicine and Microbiology, University of Iowa Carver College of Medicine, Iowa City, Iowa, United States of America, Department of Epidemiology, University of Iowa College of Public Health, Iowa City, Iowa, United States of America, VA Medical Center, Iowa City, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NE. Performed the experiments: NE. Analyzed the data: NE MW. Contributed reagents/materials/analysis tools: MW. Wrote the paper: NE MW.