About the Authors

Haotian Lin

Contributed equally to this work with: Haotian Lin, Erping Long

Roles Conceptualization, Funding acquisition, Investigation, Project administration, Writing – original draft, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Erping Long

Contributed equally to this work with: Haotian Lin, Erping Long

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Resources, Software, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Xiaohu Ding

Roles Methodology, Validation, Visualization, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Hongxing Diao

Roles Project administration, Resources, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Zicong Chen

Roles Formal analysis, Methodology, Software, Validation, Writing – review & editing

Affiliation School of Public Health, Sun Yat-sen University, Guangzhou, China

Runzhong Liu

Roles Data curation, Investigation, Methodology, Software, Writing – review & editing

Affiliation School of Public Health, Sun Yat-sen University, Guangzhou, China

Jialing Huang

Roles Formal analysis, Methodology, Validation, Writing – review & editing

Affiliation School of Public Health, Sun Yat-sen University, Guangzhou, China

Jingheng Cai

Roles Methodology, Writing – review & editing

Affiliation School of Mathematics, Sun Yat-sen University, Guangzhou, China

Shuangjuan Xu

Roles Data curation, Methodology, Writing – review & editing

Affiliation School of Mathematics, Sun Yat-sen University, Guangzhou, China

Xiayin Zhang

Roles Formal analysis, Investigation, Supervision, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Dongni Wang

Roles Project administration, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Kexin Chen

Roles Data curation, Resources, Writing – review & editing

Affiliation Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, China

Tongyong Yu

Roles Methodology, Writing – review & editing

Affiliation Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, China

Dongxuan Wu

Roles Resources, Writing – review & editing

Affiliation Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, China

Xutu Zhao

Roles Methodology, Writing – review & editing

Affiliation Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, China

Zhenzhen Liu

Roles Supervision, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Xiaohang Wu

Roles Supervision, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Yuzhen Jiang

Roles Supervision, Writing – review & editing

Affiliations State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China, UCL Institute of Ophthalmology, University College London and Moorfields Eye Hospital, London, United Kingdom

Xiao Yang

Roles Resources, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Dongmei Cui

Roles Supervision, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Wenyan Liu

Roles Resources, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Yingfeng Zheng

Roles Supervision, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Lixia Luo

Roles Supervision, Writing – review & editing

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Haibo Wang

Roles Supervision, Writing – review & editing

Affiliation First Affiliated Hospital of Sun Yat-sen University, Guangzhou, China

Chi-Chao Chan

Roles Supervision, Writing – review & editing

Affiliations State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China, Laboratory of Immunology, National Eye Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Ian G. Morgan

Roles Conceptualization, Supervision, Writing – review & editing

Affiliations State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China, ARC Centre of Excellence in Vision Science, Research School of Biology, College of Medicine, Biology and Environment, Australian National University, Canberra, Australian Capital Territory, Australia

Mingguang He

Roles Conceptualization, Data curation, Investigation, Project administration, Resources, Supervision, Writing – original draft, Writing – review & editing

Affiliations State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China, Centre for Eye Research Australia, University of Melbourne, Royal Victorian Eye and Ear Hospital, East Melbourne, Victoria, Australia

Yizhi Liu

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Project administration, Supervision, Validation, Writing – review & editing

yizhi_liu@aliyun.com (YL)

Affiliation State Key Laboratory of Ophthalmology, Clinical Research Center for Ocular Disease, Zhongshan Ophthalmic Centre, Sun Yat-sen University, Guangzhou, China

Competing Interests

The authors have no conflicts of interest to declare.