About the Authors

Anthony Sinadinos

Affiliation Molecular Medicine, School of Pharmacy and Biomedical Sciences, University of Portsmouth, Portsmouth, United Kingdom

Christopher N. J. Young

Contributed equally to this work with: Christopher N. J. Young, Rasha Al-Khalidi, Léonore Floriot, Tiphaine Henry

Affiliation Molecular Medicine, School of Pharmacy and Biomedical Sciences, University of Portsmouth, Portsmouth, United Kingdom

Rasha Al-Khalidi

Contributed equally to this work with: Christopher N. J. Young, Rasha Al-Khalidi, Léonore Floriot, Tiphaine Henry

Affiliation Molecular Medicine, School of Pharmacy and Biomedical Sciences, University of Portsmouth, Portsmouth, United Kingdom

Anna Teti

Affiliation Department of Biotechnological and Applied Clinical Sciences, University of L’Aquila, L’Aquila, Italy

Paweł Kalinski

Affiliation Departments of Surgery, Immunology, and Bioengineering, School of Medicine, University of Pittsburgh, Pittsburg, Pennsylvania, United States of America

Shafini Mohamad

Affiliation School of Engineering, University of Portsmouth, Portsmouth, United Kingdom

Léonore Floriot

Contributed equally to this work with: Christopher N. J. Young, Rasha Al-Khalidi, Léonore Floriot, Tiphaine Henry

Affiliation Platform of Behavioural Analysis (SCAC), University of Rouen, Mont-Saint-Aignan,Rouen, France

Tiphaine Henry

Contributed equally to this work with: Christopher N. J. Young, Rasha Al-Khalidi, Léonore Floriot, Tiphaine Henry

Affiliation Platform of Behavioural Analysis (SCAC), University of Rouen, Mont-Saint-Aignan,Rouen, France

Gianluca Tozzi

Affiliation School of Engineering, University of Portsmouth, Portsmouth, United Kingdom

Taiwen Jiang

Affiliation Molecular Medicine, School of Pharmacy and Biomedical Sciences, University of Portsmouth, Portsmouth, United Kingdom

Olivier Wurtz

Affiliation INSERM U982, Plate-Forme d’Imagerie PRIMACEN, IRIB, University of Rouen, Mont-Saint-Aignan, France

Alexis Lefebvre

Affiliation Platform of Behavioural Analysis (SCAC), University of Rouen, Mont-Saint-Aignan,Rouen, France

Mikhail Shugay

Affiliation Genomics of Adaptive Immunity Lab, Shemyakin and Ovchinnikov Institute of Bioorganic Chemistry and Pirogov Russian National Research Medical University, Moscow, Russia

Jie Tong

Affiliation School of Engineering, University of Portsmouth, Portsmouth, United Kingdom

David Vaudry

Affiliation INSERM U982, Plate-Forme d’Imagerie PRIMACEN, IRIB, University of Rouen, Mont-Saint-Aignan, France

Stephen Arkle

Affiliation Molecular Medicine, School of Pharmacy and Biomedical Sciences, University of Portsmouth, Portsmouth, United Kingdom

Jean-Claude doRego

Affiliations Platform of Behavioural Analysis (SCAC), University of Rouen, Mont-Saint-Aignan,Rouen, France, National Center of Scientific Research (CNRS), Caen, France

Dariusz C. Górecki

darek.gorecki@port.ac.uk

Affiliation Molecular Medicine, School of Pharmacy and Biomedical Sciences, University of Portsmouth, Portsmouth, United Kingdom

Competing Interests

The authors have declared that no conflict of interest exists.

Author Contributions

Conceived and designed the experiments: DCG J-CdR JT PK. Performed the experiments: AS CY RA-K AT SM LF TH GT TJ OW AL MS J-CdR DCG. Analyzed the data: AS CY RA-K AT PK SM LF TH GT TJ OW AL JT MS SA DV J-CdR DCG. Contributed reagents/materials/analysis tools: AT JT DV J-CdR PK MS. Wrote the first draft of the manuscript: AS J-CdR DCG. Contributed to the writing of the manuscript: AS CY RA-K AT PK SM LF TH GT TJ OW AL JT MS SA DV J-CdR DCG. Agree with the manuscript’s results and conclusions: AS CY RA-K AT PK SM LF TH GT TJ OW AL JT MS SA DV J-CdR DCG. All authors have read, and confirm that they meet, ICMJE criteria for authorship.