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Research Article
Contributed equally to this work with: Guangju Zhai, Alexander Teumer, Lisette Stolk, John R. B. Perry, Liesbeth Vandenput, Andrea D. Coviello, Annemarie Koster
Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom
⨯Contributed equally to this work with: Guangju Zhai, Alexander Teumer, Lisette Stolk, John R. B. Perry, Liesbeth Vandenput, Andrea D. Coviello, Annemarie Koster
Affiliation Interfaculty Institute for Genetics and Functional Genomics, University of Greifswald, Greifswald, Germany
⨯Contributed equally to this work with: Guangju Zhai, Alexander Teumer, Lisette Stolk, John R. B. Perry, Liesbeth Vandenput, Andrea D. Coviello, Annemarie Koster
Affiliations Department of Internal Medicine, Erasmus MC Rotterdam, Rotterdam, The Netherlands, Netherlands Consortium of Healthy Ageing, Rotterdam, The Netherlands
⨯Contributed equally to this work with: Guangju Zhai, Alexander Teumer, Lisette Stolk, John R. B. Perry, Liesbeth Vandenput, Andrea D. Coviello, Annemarie Koster
Affiliations Genetics of Complex Traits, Peninsula Medical School, University of Exeter, Exeter, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom
⨯Contributed equally to this work with: Guangju Zhai, Alexander Teumer, Lisette Stolk, John R. B. Perry, Liesbeth Vandenput, Andrea D. Coviello, Annemarie Koster
Affiliation Department of Internal Medicine, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden
⨯Contributed equally to this work with: Guangju Zhai, Alexander Teumer, Lisette Stolk, John R. B. Perry, Liesbeth Vandenput, Andrea D. Coviello, Annemarie Koster
Affiliation Sections of General Internal Medicine, Preventive Medicine, Cardiology and Endocrinology, Diabetes and Nutrition, Department of Medicine, Boston University School of Medicine, Boston, Massachusetts, United States of America
⨯Contributed equally to this work with: Guangju Zhai, Alexander Teumer, Lisette Stolk, John R. B. Perry, Liesbeth Vandenput, Andrea D. Coviello, Annemarie Koster
Affiliation Laboratory for Epidemiology, Demography, and Biometry, National Institute on Aging, Bethesda, Maryland, United States of America
⨯Affiliations Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom
⨯Affiliation Section of Endocrinology, Diabetes, and Nutrition, Claude D. Pepper Older Americans Independence Center, Boston University School of Medicine, Boston, Massachusetts, United States of America
⨯Affiliation Department of Internal Medicine, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden
⨯Affiliation Department of Internal Medicine, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden
⨯Affiliation Interfaculty Institute for Genetics and Functional Genomics, University of Greifswald, Greifswald, Germany
⨯Affiliation Clinical Research Branch, National Institute on Aging, Baltimore, Maryland, United States of America
⨯Affiliation Genetics of Complex Traits, Peninsula Medical School, University of Exeter, Exeter, United Kingdom
⨯Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom
⨯Affiliations Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom, Wellcome Trust Sanger Institute, Hixton, United Kingdom
⨯Affiliation Institute for Clinical Chemistry and Laboratory Medicine, University of Greifswald, Greifswald, Germany
⨯Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom
⨯Affiliations Netherlands Consortium of Healthy Ageing, Rotterdam, The Netherlands, Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands
⨯Affiliation Hebrew Senior Life Institute for Aging Research and Harvard Medical School, Boston, Massachusetts, United States of America
⨯Affiliation Center of Pharmacology and Experimental Therapeutics, Department of Pharmacology, University of Greifswald, Greifswald, Germany
⨯Affiliation Department of Epidemiology and Prevention, Wake Forest University Health Sciences, Winston-Salem, North Carolina, United States of America
⨯Affiliation Department of Biostatistics, Boston University School of Public Health, Boston, Massachusetts, United States of America
⨯Affiliation Department of Internal Medicine and Biomedical Sciences, Section of Geriatrics, University of Parma, Parma, Italy
⨯Affiliation Department of Internal Medicine, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden
⨯Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom
⨯Affiliation Genetics of Complex Traits, Peninsula Medical School, University of Exeter, Exeter, United Kingdom
⨯Affiliation Department of Epidemiology, University of Pittsburgh, Pittsburg, Pennsylvania, United States of America
⨯Affiliations Wellcome Trust Sanger Institute, Hixton, United Kingdom, Department of Genetic Medicine and Development, University of Geneva Medical School, Geneva, Switzerland
⨯Affiliation Department of Psychiatry, VU University Medical Center, Amsterdam, The Netherlands
⨯Affiliations Sections of General Internal Medicine, Preventive Medicine, Cardiology and Endocrinology, Diabetes and Nutrition, Department of Medicine, Boston University School of Medicine, Boston, Massachusetts, United States of America, The National Heart Lung and Blood Institute's Framingham Heart Study, Framingham, Massachusetts, United States of America
⨯Affiliations Department of Internal Medicine, Erasmus MC Rotterdam, Rotterdam, The Netherlands, Netherlands Consortium of Healthy Ageing, Rotterdam, The Netherlands
⨯Affiliations Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom, Wellcome Trust Sanger Institute, Hixton, United Kingdom
⨯Affiliations Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom, Wellcome Trust Sanger Institute, Hixton, United Kingdom
⨯Affiliations Department of Internal Medicine, Erasmus MC Rotterdam, Rotterdam, The Netherlands, Netherlands Consortium of Healthy Ageing, Rotterdam, The Netherlands, Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands
⨯Affiliation Institute for Community Medicine, University of Greifswald, Greifswald, Germany
⨯Affiliations Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom, Department of Endocrinology and Diabetes, Sir Charles Gairdner Hospital, Nedlands, Australia, School of Medicine and Pharmacology, University of Western Australia, Nedlands, Australia
⨯Affiliation Molecular Medicine – Computational Biology, Pfizer Worldwide R&D, Groton, Connecticut, United States of America
⨯Affiliation Department of Biostatistics, Boston University School of Public Health, Boston, Massachusetts, United States of America
⨯¶These senior authors were joint senior authors on this work.
Affiliation Laboratory for Epidemiology, Demography, and Biometry, National Institute on Aging, Bethesda, Maryland, United States of America
⨯¶These senior authors were joint senior authors on this work.
Affiliations Sections of General Internal Medicine, Preventive Medicine, Cardiology and Endocrinology, Diabetes and Nutrition, Department of Medicine, Boston University School of Medicine, Boston, Massachusetts, United States of America, The National Heart Lung and Blood Institute's Framingham Heart Study, Framingham, Massachusetts, United States of America
⨯¶These senior authors were joint senior authors on this work.
Affiliation Department of Internal Medicine, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden
⨯¶These senior authors were joint senior authors on this work.
Affiliation Genetics of Complex Traits, Peninsula Medical School, University of Exeter, Exeter, United Kingdom
⨯¶These senior authors were joint senior authors on this work.
Affiliation Department of Internal Medicine, Erasmus MC Rotterdam, Rotterdam, The Netherlands
⨯* E-mail: tim.spector@kcl.ac.uk
Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom
⨯¶These senior authors were joint senior authors on this work.
Affiliation Institute for Clinical Chemistry and Laboratory Medicine, University of Greifswald, Greifswald, Germany
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