About the Authors

Ying Xiong

Contributed equally to this work with: Ying Xiong, Liqing Fan

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Project administration, Resources, Software, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China

Liqing Fan

Contributed equally to this work with: Ying Xiong, Liqing Fan

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Resources, Software, Visualization, Writing – original draft, Writing – review & editing

Affiliations National Forest Ecosystem Observation & Research Station of Nyingchi Tibet, Institute of Plateau Ecology, Tibet Agriculture & Animal Husbandry University, Linzhi City, China, Key Laboratory of Forest Ecology in Tibet Plateau (Tibet Agriculture & Animal Husbandry University), Ministry of Education, Linzhi City, China

Yan Hao

Roles Formal analysis, Methodology, Software, Writing – review & editing

Affiliations Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China

Yalin Cheng

Roles Formal analysis, Methodology, Writing – review & editing

Affiliations Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China

Yongbin Chang

Roles Formal analysis, Methodology, Writing – review & editing

Affiliations Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China

Jing Wang

Roles Formal analysis, Writing – review & editing

Affiliations Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China

Haiyan Lin

Roles Writing – review & editing

Affiliation Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Gang Song

Roles Methodology, Resources, Writing – review & editing

Affiliation Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Yanhua Qu

Roles Methodology, Resources, Writing – review & editing

Affiliation Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Fumin Lei

Roles Conceptualization, Funding acquisition, Investigation, Project administration, Resources, Supervision, Validation, Writing – original draft, Writing – review & editing

leifm@ioz.ac.cn

Affiliations Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China, Center for Excellence in Animal Evolution and Genetics, Chinese Academy of Sciences, Kunming, China

Competing Interests

The authors have declared that no competing interests exist.