About the Authors

Xiaoxi Liu

Roles Conceptualization, Formal analysis, Investigation, Methodology, Software, Visualization, Writing – original draft, Writing – review & editing

Affiliation Genome Immunobiology RIKEN Hakubi Research Team, RIKEN Cluster for Pioneering Research and RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Shunichi Kosugi

Roles Data curation, Investigation, Methodology, Software

Affiliation Laboratory for Statistical and Translational Genetics, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Rie Koide

Roles Investigation, Methodology

Affiliation Genome Immunobiology RIKEN Hakubi Research Team, RIKEN Cluster for Pioneering Research and RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Yoshiki Kawamura

Roles Investigation, Resources

Affiliation Department of Pediatrics, Fujita Health University School of Medicine, Toyoake, Japan

Jumpei Ito

Roles Validation

Affiliation Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Hiroki Miura

Roles Investigation, Resources

Affiliation Department of Pediatrics, Fujita Health University School of Medicine, Toyoake, Japan

Nana Matoba

Roles Data curation, Software

Current address: Department of Genetics and UNC Neuroscience Center, University of North Carolina at Chapel Hill, United States of America.

Affiliation Laboratory for Statistical and Translational Genetics, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Motomichi Matsuzaki

Roles Validation

Affiliation Statistical Genetics Team, RIKEN Center for Advanced Intelligence Project, Tokyo, Japan

Masashi Fujita

Roles Validation

Affiliation Laboratory for Cancer Genomics, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Anselmo Jiro Kamada

Roles Resources

Affiliation Genome Immunobiology RIKEN Hakubi Research Team, RIKEN Cluster for Pioneering Research and RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Hidewaki Nakagawa

Roles Supervision, Validation

Affiliation Laboratory for Cancer Genomics, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Gen Tamiya

Roles Supervision, Validation

Affiliation Statistical Genetics Team, RIKEN Center for Advanced Intelligence Project, Tokyo, Japan

Koichi Matsuda

Roles Funding acquisition, Project administration, Resources

Affiliations Laboratory of Molecular Medicine, Institute of Medical Science, The University of Tokyo, Tokyo, Japan, Laboratory for Clinical Genome Sequencing, Graduate School of Frontier Sciences, The University of Tokyo, Tokyo, Japan

Yoshinori Murakami

Roles Funding acquisition, Project administration, Resources

Affiliation Division of Molecular Pathology, Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Michiaki Kubo

Roles Funding acquisition, Project administration, Resources

Affiliation RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Amr Aswad

Roles Writing – review & editing

Affiliation Institut für Virologie, Freie Universität Berlin, Berlin, Germany

Kei Sato

Roles Supervision, Validation

Affiliation Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Yukihide Momozawa

Roles Data curation, Funding acquisition, Project administration, Resources, Supervision

Affiliation Laboratory for Genotyping Development, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Jun Ohashi

Roles Formal analysis, Writing – original draft, Writing – review & editing

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan

Chikashi Terao

Roles Data curation, Funding acquisition, Project administration, Resources, Supervision, Writing – review & editing

Affiliation Laboratory for Statistical and Translational Genetics, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Tetsushi Yoshikawa

Roles Funding acquisition, Resources, Supervision

Affiliation Department of Pediatrics, Fujita Health University School of Medicine, Toyoake, Japan

Nicholas F. Parrish

Roles Conceptualization, Formal analysis, Funding acquisition, Project administration, Supervision, Visualization, Writing – original draft, Writing – review & editing

nicholas.parrish@riken.jp

Affiliation Genome Immunobiology RIKEN Hakubi Research Team, RIKEN Cluster for Pioneering Research and RIKEN Center for Integrative Medical Sciences, Yokohama, Japan

Yoichiro Kamatani

Roles Data curation, Funding acquisition, Project administration, Resources, Supervision, Writing – review & editing

Affiliations Laboratory for Statistical and Translational Genetics, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan, Laboratory of Complex Trait Genomics, Graduate School of Frontier Sciences, The University of Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.