About the Authors

Elliott W. Abrams

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America, Department of Biology, Purchase College, The State University of New York, Purchase, New York, United States of America

Ricardo Fuentes

Roles Formal analysis, Investigation, Methodology, Validation, Visualization, Writing – review & editing

Current address: Department of Cell Biology, Faculty of Biological Sciences, University of Concepcion, Concepcion, Chile

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Florence L. Marlow

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Supervision, Writing – review & editing

Current address: Department of Cell, Developmental and Regenerative Biology, Icahn School of Medicine Mount Sinai, New York, New York, United States of America

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Manami Kobayashi

Roles Formal analysis, Investigation, Methodology, Validation, Visualization, Writing – review & editing

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Hong Zhang

Roles Conceptualization, Formal analysis, Investigation, Methodology

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Sumei Lu

Roles Formal analysis, Investigation, Methodology

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Lee Kapp

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Shai R. Joseph

Roles Conceptualization, Formal analysis, Investigation, Methodology

Affiliation Max Planck Institute of Molecular Cell Biology and Genetics, Dresden, Germany

Amy Kugath

Roles Formal analysis, Investigation, Methodology

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Tripti Gupta

Roles Conceptualization, Formal analysis, Investigation, Methodology, Supervision

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Virginia Lemon

Roles Formal analysis, Investigation, Methodology

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Greg Runke

Roles Formal analysis, Investigation, Methodology

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Amanda A. Amodeo

Roles Conceptualization, Formal analysis, Investigation, Methodology, Writing – review & editing

Affiliation Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America

Nadine L. Vastenhouw

Roles Conceptualization, Formal analysis, Investigation, Methodology, Supervision, Writing – review & editing

Affiliation Max Planck Institute of Molecular Cell Biology and Genetics, Dresden, Germany

Mary C. Mullins

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Supervision, Visualization, Writing – review & editing

mullins@pennmedicine.upenn.edu

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.