About the Authors

Kati J. Dillard

Roles Conceptualization, Formal analysis, Investigation, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, Department of Medical and Clinical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Research Center, Helsinki, Finland, Veterinary Bacteriology and Pathology Research Unit, Finnish Food Authority, Helsinki, Finland

Matthias Ochs

Roles Formal analysis, Writing – original draft, Writing – review & editing

Affiliations Institute of Functional and Applied Anatomy, Hannover Medical School, Hannover, Germany, Institute of Functional Anatomy, Charité - Universitaetsmedizin Berlin, Berlin, Germany, German Center for Lung Research (DZL), Berlin, Germany

Julia E. Niskanen

Roles Investigation, Visualization, Writing – review & editing

Affiliations Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, Department of Medical and Clinical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Research Center, Helsinki, Finland

Meharji Arumilli

Roles Data curation, Resources, Software, Writing – review & editing

Affiliations Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, Department of Medical and Clinical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Research Center, Helsinki, Finland

Jonas Donner

Roles Formal analysis, Resources, Validation, Writing – review & editing

Affiliation Genoscoper Laboratories Ltd (Wisdom Health), Helsinki, Finland

Kaisa Kyöstilä

Roles Formal analysis, Validation

Affiliations Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, Department of Medical and Clinical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Research Center, Helsinki, Finland

Marjo K. Hytönen

Contributed equally to this work with: Marjo K. Hytönen, Marjukka Anttila, Hannes Lohi

Roles Conceptualization, Formal analysis, Investigation, Project administration, Supervision, Validation, Writing – review & editing

‡ These authors are joint senior authors on this work

Affiliations Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, Department of Medical and Clinical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Research Center, Helsinki, Finland

Marjukka Anttila

Contributed equally to this work with: Marjo K. Hytönen, Marjukka Anttila, Hannes Lohi

Roles Conceptualization, Formal analysis, Investigation, Project administration, Resources, Supervision, Validation, Writing – original draft, Writing – review & editing

‡ These authors are joint senior authors on this work

Affiliation Veterinary Bacteriology and Pathology Research Unit, Finnish Food Authority, Helsinki, Finland

Hannes Lohi

Contributed equally to this work with: Marjo K. Hytönen, Marjukka Anttila, Hannes Lohi

Roles Conceptualization, Funding acquisition, Project administration, Resources, Supervision, Validation, Writing – original draft, Writing – review & editing

hannes.lohi@helsinki.fi

‡ These authors are joint senior authors on this work

Affiliations Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, Department of Medical and Clinical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Research Center, Helsinki, Finland

Competing Interests

I have read the journal's policy and the authors of this manuscript have the following competing interests: HL has consulted Genoscoper Laboratories Oy, which provides genetic tests for dogs, including the finding in this study. JD is an employee of Genoscoper Laboratories Oy.