About the Authors

Alexandra M. Sparks

Roles Conceptualization, Formal analysis, Investigation, Methodology, Visualization, Writing – original draft, Writing – review & editing

A.M.Sparks@leeds.ac.uk

Affiliations Institutes of Evolutionary Biology and Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom, Faculty of Biological Sciences, School of Biology, University of Leeds, Leeds, United Kingdom

Kathryn Watt

Roles Investigation, Methodology

Affiliation Institutes of Evolutionary Biology and Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Rona Sinclair

Roles Investigation

Affiliation Institutes of Evolutionary Biology and Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Jill G. Pilkington

Roles Investigation, Project administration

Affiliation Institutes of Evolutionary Biology and Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Josephine M. Pemberton

Roles Funding acquisition, Project administration, Resources, Writing – review & editing

Affiliation Institutes of Evolutionary Biology and Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Tom N. McNeilly

Roles Conceptualization, Funding acquisition, Methodology, Resources, Supervision, Writing – review & editing

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Midlothian, United Kingdom

Daniel H. Nussey

Roles Conceptualization, Funding acquisition, Methodology, Project administration, Resources, Supervision, Writing – original draft, Writing – review & editing

Affiliation Institutes of Evolutionary Biology and Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Susan E. Johnston

Roles Conceptualization, Formal analysis, Methodology, Resources, Supervision, Visualization, Writing – original draft, Writing – review & editing

Affiliation Institutes of Evolutionary Biology and Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.