About the Authors

Thomas W. Marchant

Roles Data curation, Formal analysis, Investigation, Visualization, Writing – original draft, Writing – review & editing

Affiliation The Roslin Institute and Royal (Dick) School for Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Elisabeth Dietschi

Roles Data curation, Investigation, Writing – review & editing

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Ulrich Rytz

Roles Formal analysis, Writing – review & editing

Affiliation Department of Clinical Veterinary Medicine, Division of Small Animal Surgery, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Peter Schawalder

Roles Data curation, Investigation, Writing – review & editing

Affiliation Department of Clinical Veterinary Medicine, Division of Small Animal Surgery, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Vidhya Jagannathan

Roles Data curation, Formal analysis, Methodology

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Sheida Hadji Rasouliha

Roles Investigation

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Corinne Gurtner

Roles Investigation, Writing – review & editing

Affiliation Institute for Animal Pathology, Department of Infectious Diseases and Pathobiology, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Andreas S. Waldvogel

Roles Investigation

Affiliation Institute for Animal Pathology, Department of Infectious Diseases and Pathobiology, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Ronan S. Harrington

Roles Data curation, Investigation

Affiliation The Roslin Institute and Royal (Dick) School for Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Michaela Drögemüller

Roles Data curation

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Jeffrey Kidd

Roles Investigation, Resources

Affiliation Department of Human Genetics, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Elaine A. Ostrander

Roles Resources, Writing – review & editing

Affiliation Cancer Genetics and Comparative Genomics Branch, National Human Genome Research Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Amanda Warr

Roles Investigation, Writing – review & editing

Affiliation The Roslin Institute and Royal (Dick) School for Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Mick Watson

Roles Investigation, Writing – review & editing

Affiliation The Roslin Institute and Royal (Dick) School for Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

David Argyle

Roles Supervision

Affiliation The Roslin Institute and Royal (Dick) School for Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Gert Ter Haar

Roles Resources

Current address: Specialistische Dierenkliniek Utrecht, Utrecht, Netherlands

Affiliation Department of Clinical Sciences and Services, Royal Veterinary College, Hertfordshire, United Kingdom

Dylan N. Clements

Roles Supervision, Writing – review & editing

Affiliation The Roslin Institute and Royal (Dick) School for Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Tosso Leeb

Roles Funding acquisition, Investigation, Methodology, Resources, Writing – original draft, Writing – review & editing

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Jeffrey J. Schoenebeck

Roles Conceptualization, Formal analysis, Funding acquisition, Methodology, Project administration, Supervision, Writing – original draft, Writing – review & editing

jeff.schoenebeck@roslin.ed.ac.uk

Affiliation The Roslin Institute and Royal (Dick) School for Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Competing Interests

I have read the journal’s policy and the authors of this manuscript have the following competing interests: TL is an associate editor of PLoS Genetics.