About the Authors

Pradeep Suri

Roles Conceptualization, Funding acquisition, Investigation, Methodology, Project administration, Supervision, Writing – original draft, Writing – review & editing

pradeep.suri@va.gov

Affiliations Seattle Epidemiologic Research and Information Center (ERIC), Department of Veterans Affairs Office of Research and Development, Seattle, Washington, United States of America, Division of Rehabilitation Care Services, VA Puget Sound Health Care System, Seattle, Washington, United States of America, Department of Rehabilitation Medicine, University of Washington, Seattle, Washington, United States of America

Melody R. Palmer

Roles Formal analysis, Methodology, Writing – original draft, Writing – review & editing

Affiliation Medical Genetics, Department of Medicine, University of Washington, Seattle, Washington, United States of America

Yakov A. Tsepilov

Roles Formal analysis, Methodology, Writing – review & editing

Affiliations Polyomica, ‘s-Hertogenbosch, the Netherlands, Laboratory of Theoretical and Applied Functional Genomics, Novosibirsk State University, Novosibirsk, Russia, Laboratory of Recombination and Segregation Analysis, Institute of Cytology and Genetics SD RAS, Novosibirsk, Russia

Maxim B. Freidin

Roles Formal analysis, Methodology, Writing – original draft, Writing – review & editing

Affiliation Department of Twin Research and Genetic Epidemiology, King’s College London, London, United Kingdom

Cindy G. Boer

Roles Formal analysis, Methodology, Writing – review & editing

Affiliation Department of Internal Medicine, Erasmus Medical Center, Rotterdam, the Netherlands

Michelle S. Yau

Roles Formal analysis, Writing – original draft, Writing – review & editing

Affiliations Institute for Aging Research, Hebrew SeniorLife, Boston, Massachusetts, United States of America, Department of Medicine, Beth Israel Deaconess Medical Center and Harvard Medical School, Boston, Massachusetts, United States of America

Daniel S. Evans

Roles Formal analysis, Project administration, Writing – review & editing

Affiliation California Pacific Medical Center Research Institute, San Francisco, California, United States of America

Andrea Gelemanovic

Roles Formal analysis, Writing – review & editing

Affiliation Department of Public Health, University of Split Medical School, Split, Croatia

Traci M. Bartz

Roles Formal analysis, Writing – review & editing

Affiliations Cardiovascular Health Research Unit and Department of Medicine, University of Washington, Seattle, Washington, United States of America, Department of Biostatistics, University of Washington, Seattle, Washington, United States of America

Maria Nethander

Roles Formal analysis, Writing – review & editing

Affiliation Department of Medicine, University of Göteborg, Göteborg, Sweden

Liubov Arbeeva

Roles Formal analysis, Writing – review & editing

Affiliation Department of Medicine, School of Medicine, University of North Carolina, Chapel Hill, North Carolina, United States of America

Lennart Karssen

Roles Formal analysis, Methodology, Supervision, Writing – review & editing

Affiliation Polyomica, ‘s-Hertogenbosch, the Netherlands

Tuhina Neogi

Roles Methodology, Supervision, Writing – review & editing

Affiliation Clinical Epidemiology Unit, Department of Medicine, Boston University School of Medicine, Boston, Massachusetts, United States of America

Archie Campbell

Roles Formal analysis, Project administration, Writing – review & editing

Affiliation Centre for Genomic and Experimental Medicine, MRC Institute of Genetics & Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Dan Mellstrom

Roles Funding acquisition, Methodology, Supervision, Writing – review & editing

Affiliation Geriatric Medicine, Department of Internal Medicine and Clinical Nutrition, Institute of Medicine, University of Gothenburg, Sweden

Claes Ohlsson

Roles Funding acquisition, Supervision, Writing – review & editing

Affiliation Department of Internal Medicine and Clinical Nutrition, Institute of Medicine, University of Gothenburg, Göteborg, Sweden

Lynn M. Marshall

Roles Methodology, Writing – review & editing

Affiliation Department of Orthopedics and Rehabilitation, Oregon Health and Science University, Portland, Oregon, United States of America

Eric Orwoll

Roles Funding acquisition, Supervision, Writing – review & editing

Affiliation Department of Medicine, Oregon Health and Science University, Portland, Oregon, United States of America

Andre Uitterlinden

Roles Funding acquisition, Supervision, Writing – review & editing

Affiliation Department of Internal Medicine, Erasmus Medical Center, Rotterdam, the Netherlands

Jerome I. Rotter

Roles Investigation, Writing – review & editing

Affiliations Institute for Translational Genomics and Population Sciences, Los Angeles Biomedical Research Institute, Harbor-UCLA Medical Center, Torrance, California, United States of America, Division of Genomic Outcomes, Departments of Pediatrics and Medicine, Harbor-UCLA Medical Center, Torrance, California, United States of America

Gordan Lauc

Roles Supervision, Writing – review & editing

Affiliations Genos Ltd, Osijek, Croatia, Faculty of Pharmacy and Biochemistry, University of Zagreb, Zagreb, Croatia

Bruce M. Psaty

Roles Funding acquisition, Methodology, Supervision, Writing – review & editing

Affiliations Cardiovascular Health Research Unit and Department of Medicine, University of Washington, Seattle, Washington, United States of America, Department of Health Services, University of Washington, Seattle, Washington, United States of America, Department of Epidemiology, University of Washington, Seattle, Washington, United States of America, Kaiser Permanente Washington Health Research Institute, Kaiser Permanente Washington, Seattle, United States of America

Magnus K. Karlsson

Roles Funding acquisition, Supervision, Writing – review & editing

Affiliation Department of Orthopedics, Skane University Hospital, Lund University, Malmö, Sweden

Nancy E. Lane

Roles Funding acquisition, Supervision, Writing – review & editing

Affiliation Departments of Medicine and Rheumatology, University of California Davis, Sacramento, California, United States of America

Gail P. Jarvik

Roles Methodology, Supervision, Writing – review & editing

Affiliations Medical Genetics, Department of Medicine, University of Washington, Seattle, Washington, United States of America, Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Ozren Polasek

Roles Funding acquisition, Project administration, Supervision, Writing – review & editing

Affiliations Department of Public Health, University of Split Medical School, Split, Croatia, Hospital “Sveti Ivan”, Zagreb, Croatia

Marc Hochberg

Roles Methodology, Supervision, Writing – review & editing

Affiliation Departments of Medicine and Epidemiology, University of Maryland, Baltimore, Maryland, United States of America

Joanne M. Jordan

Roles Methodology, Supervision, Writing – review & editing

Affiliation Department of Medicine, School of Medicine, University of North Carolina, Chapel Hill, North Carolina, United States of America

Joyce B. J. Van Meurs

Roles Project administration, Supervision, Writing – review & editing

Affiliation Department of Internal Medicine, Erasmus Medical Center, Rotterdam, the Netherlands

Rebecca Jackson

Roles Methodology, Writing – review & editing

Affiliation Department of Medicine, The Ohio State University, Columbus, Ohio, United States of America

Carrie M. Nielson

Roles Methodology, Writing – review & editing

Affiliation School of Public Health, Oregon Health and Science University, Portland, Oregon, United States of America

Braxton D. Mitchell

Roles Methodology, Writing – review & editing

Affiliations Departments of Medicine and Epidemiology, University of Maryland, Baltimore, Maryland, United States of America, Geriatric Research, Education and Clinical Center, Veterans Affairs Medical Center, Baltimore, Maryland, United States of America

Blair H. Smith

Roles Methodology, Supervision, Writing – review & editing

Affiliation Division of Population Health Sciences, School of Medicine, University of Dundee, Dundee, United Kingdom

Caroline Hayward

Roles Formal analysis, Methodology, Writing – review & editing

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics & Molecular Medicine, University of Edinburgh, United Kingdom

Nicholas L. Smith

Roles Conceptualization, Methodology, Writing – original draft, Writing – review & editing

Affiliations Seattle Epidemiologic Research and Information Center (ERIC), Department of Veterans Affairs Office of Research and Development, Seattle, Washington, United States of America, Department of Epidemiology, University of Washington, Seattle, Washington, United States of America, Kaiser Permanente Washington Health Research Institute, Kaiser Permanente Washington, Seattle, United States of America

Yurii S. Aulchenko

Roles Conceptualization, Formal analysis, Methodology, Supervision, Writing – original draft, Writing – review & editing

Affiliation Polyomica, ‘s-Hertogenbosch, the Netherlands

Frances M. K. Williams

Roles Conceptualization, Methodology, Supervision, Writing – original draft, Writing – review & editing

Affiliation Department of Twin Research and Genetic Epidemiology, King’s College London, London, United Kingdom

Competing Interests

BMP serves on the DSMB of a clinical trial funded by Zoll LifeCor and on the Steering Committee of the Yale Open Data Access Project funded by Johnson & Johnson; YSA and LK are founders and co-owners of PolyOmica, a private genomics research organization; these relationships do not pose known conflicts with the content of this work presented. We the authors do not have any other financial interests that could create a potential conflict or the appearance of a potential conflict of interest with regard to this work.