About the Authors

Shin-ichi Tokuhiro

Contributed equally to this work with: Shin-ichi Tokuhiro, Miki Tokuoka, Kenji Kobayashi

Affiliation Department of Zoology, Graduate School of Science, Kyoto University, Kyoto, Japan

Miki Tokuoka

Contributed equally to this work with: Shin-ichi Tokuhiro, Miki Tokuoka, Kenji Kobayashi

Affiliation Department of Zoology, Graduate School of Science, Kyoto University, Kyoto, Japan

Kenji Kobayashi

Contributed equally to this work with: Shin-ichi Tokuhiro, Miki Tokuoka, Kenji Kobayashi

Affiliation Department of Zoology, Graduate School of Science, Kyoto University, Kyoto, Japan

Atsushi Kubo

Affiliation Department of Zoology, Graduate School of Science, Kyoto University, Kyoto, Japan

Izumi Oda-Ishii

Affiliation Department of Zoology, Graduate School of Science, Kyoto University, Kyoto, Japan

Yutaka Satou

yutaka@ascidian.zool.kyoto-u.ac.jp

Affiliation Department of Zoology, Graduate School of Science, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: ST MT KK YS. Data curation: YS. Formal analysis: KK YS. Funding acquisition: YS. Investigation: ST MT KK AK IOI. Methodology: AK IOI. Project administration: YS. Resources: ST MT KK AK IO YS. Supervision: YS. Validation: ST MT KK AK IOI YS. Visualization: ST MT KK YS. Writing – original draft: YS. Writing – review & editing: ST MT KK AK IOI YS.