About the Authors

Tian Ge

tge1@mgh.harvard.edu (TG); jsmoller@hms.harvard.edu (JWS)

Affiliations Athinoula A. Martinos Center for Biomedical Imaging, Massachusetts General Hospital / Harvard Medical School, Charlestown, MA, United States of America, Psychiatric and Neurodevelopmental Genetics Unit, Center for Genomic Medicine, Massachusetts General Hospital, Boston, MA, United States of America, Stanley Center for Psychiatric Research, Broad Institute of MIT and Harvard, Cambridge, MA, United States of America

Chia-Yen Chen

Affiliations Psychiatric and Neurodevelopmental Genetics Unit, Center for Genomic Medicine, Massachusetts General Hospital, Boston, MA, United States of America, Stanley Center for Psychiatric Research, Broad Institute of MIT and Harvard, Cambridge, MA, United States of America, Analytic and Translational Genetics Unit, Center for Genomic Medicine, Massachusetts General Hospital, Boston, MA, United States of America

Benjamin M. Neale

Affiliations Psychiatric and Neurodevelopmental Genetics Unit, Center for Genomic Medicine, Massachusetts General Hospital, Boston, MA, United States of America, Stanley Center for Psychiatric Research, Broad Institute of MIT and Harvard, Cambridge, MA, United States of America, Analytic and Translational Genetics Unit, Center for Genomic Medicine, Massachusetts General Hospital, Boston, MA, United States of America

Mert R. Sabuncu

Contributed equally to this work with: Mert R. Sabuncu, Jordan W. Smoller

Affiliations Athinoula A. Martinos Center for Biomedical Imaging, Massachusetts General Hospital / Harvard Medical School, Charlestown, MA, United States of America, School of Electrical and Computer Engineering and Meinig School of Biomedical Engineering, Cornell University, Ithaca, NY, United States of America

Jordan W. Smoller

Contributed equally to this work with: Mert R. Sabuncu, Jordan W. Smoller

tge1@mgh.harvard.edu (TG); jsmoller@hms.harvard.edu (JWS)

Affiliations Psychiatric and Neurodevelopmental Genetics Unit, Center for Genomic Medicine, Massachusetts General Hospital, Boston, MA, United States of America, Stanley Center for Psychiatric Research, Broad Institute of MIT and Harvard, Cambridge, MA, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: TG MRS JWS. Data curation: TG JWS. Formal analysis: TG CYC. Funding acquisition: JWS MRS. Methodology: TG CYC MRS JWS. Resources: MRS JWS. Software: TG CYC. Supervision: JWS MRS. Validation: TG. Visualization: TG. Writing – original draft: TG JWS. Writing – review & editing: TG CYC BMN MRS JWS.