About the Authors

Gang Li

gli11@partners.org (GL); pnigrovic@partners.org (PAN)

Affiliation Division of Rheumatology, Immunology and Allergy, Brigham and Women’s Hospital, Boston, Massachusetts, United States of America

Pierre Cunin

Affiliation Division of Rheumatology, Immunology and Allergy, Brigham and Women’s Hospital, Boston, Massachusetts, United States of America

Di Wu

Current address: Department of Periodontology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Affiliations Division of Rheumatology, Immunology and Allergy, Brigham and Women’s Hospital, Boston, Massachusetts, United States of America, Department of Statistics, Harvard University, Cambridge, Massachusetts, United States of America, Centre for Cancer Research, Monash Institute of Medical Research, Monash University, Clayton, Victoria, Australia

Dorothée Diogo

Current address: Merck Research Laboratories, Boston, Massachusetts, United States of America

Affiliations Division of Rheumatology, Immunology and Allergy, Brigham and Women’s Hospital, Boston, Massachusetts, United States of America, Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute, Cambridge, Massachusetts, United States of America

Yu Yang

Affiliation Division of Rheumatology, Immunology and Allergy, Brigham and Women’s Hospital, Boston, Massachusetts, United States of America

Yukinori Okada

Affiliations Department of Human Genetics and Disease Diversity, Graduate School of Medical and Dental Sciences, Tokyo Medical and Dental University, Tokyo, Japan, Laboratory for Statistical Analysis, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan, Department of Statistical Genetics, Osaka University Graduate School of Medicine, Osaka, Japan

Robert M. Plenge

Current address: Merck Research Laboratories, Boston, Massachusetts, United States of America

Affiliation Division of Rheumatology, Immunology and Allergy, Brigham and Women’s Hospital, Boston, Massachusetts, United States of America

Peter A. Nigrovic

gli11@partners.org (GL); pnigrovic@partners.org (PAN)

Affiliations Division of Rheumatology, Immunology and Allergy, Brigham and Women’s Hospital, Boston, Massachusetts, United States of America, Division of Immunology, Boston Children’s Hospital, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: GL RMP PAN. Formal analysis: GL DW DD YO. Funding acquisition: GL PAN. Investigation: GL PC YY. Methodology: GL. Project administration: GL PAN. Resources: GL RMP PAN. Supervision: GL RMP PAN. Visualization: GL. Writing – original draft: GL PAN. Writing – review & editing: GL PC DW DD YY YO RMP PAN.